450 Wilson —Spermatogenesis in the Bryophyta. 
‘ Nebenkorper 5 forms a part of the body of the spermatozoid. In a recent 
communication Yamanouchi (76) has described a body in the spermatid of 
Nephrodium which arises near the nucleus, but the origin of which was not 
ascertained. This body, which he designates the c Nebenkern is found per- 
sisting in the vesicle of the mature spermatozoid. 
In the present investigation two bodies have been constantly discovered 
in the spermatids, and these in the three plants examined take no direct 
part in the formation of the spermatozoid, but are found ultimately in the 
vesicle. The limosphere, as already shown, is easily distinguished in all 
cases by its peculiar form, and no doubt corresponds to the ‘ Nebenkorper ’ 
described in P oly trichum by J. and W. Docters van Leeuwen-Reijnvaan 
(41) and by Arens (2) in Mnium hornum . Up to the present no such 
structure has been discovered in the Hepaticae. The accessory body is 
probably similar to the body described by J. and W. Docters van Leeuwen- 
Reijnvaan in Polytrichum produced by the third constriction of the nucleus. 
With this exception it has not yet been recorded in the Bryophyta or else- 
where. At present it cannot be stated which of these two bodies is equiva- 
lent to the ‘ Nebenkorper 5 described by Ikeno in Marchantia. 
There seems to be no doubt that in Atrichum undulatum and Mnium 
honiitm both the limosphere and the accessory body are derived either 
directly or indirectly from the nucleus, and although in the case of Pellia 
their origin has not been determined, it seems probable that here, too, they 
are derived from the nucleus. 
The occurrence of structures corresponding to the rod-like bodies dis- 
covered in Mnium hornum has not, up to the present, been recorded in the 
spermatids of plants. Although these structures have not been found in 
Atrichum , it is still possible that they exist in this plant and that their 
occurrence has been overlooked on account of the short period during which 
they are present. Somewhat similar structures have been discovered in 
the spermatocytes and spermatids of a considerable number of animals, and 
these are variously referred to as chromidia, chondromites, mitochondria, 
pseudochromosomes, &c. 
It would be out of place here to attempt even a partial summary of 
the results obtained in animals in this connexion, but a few will be men- 
tioned. Gross (26) in Pyrrhocoris describes pseudochromosomes in sper- 
matocytes I, II, and the spermatids. In the latter these give rise to the 
‘Nebenkern’, which is somewhat ring-like in form. Schreiner (59) states 
that chondromites and chromatic bodies are present during the spermato- 
genesis of Myxine . Oettinger (48) describes the occurrence of mitochondria 
in the spermatids of some Myriapods. Some of these are ‘ fadenformige 
Mitochondrien ’, but others exist in the form of a sphere, which at first 
possesses a dark -staining periphery. Popoff (49) in Helix and Paludina 
states that chromidia are formed from granules extruded from the nucleus 
