Studies in some East Indian Hepaticae. 
Calobryum Blumei, N. ab E. 
BY 
D. H. CAMPBELL. 
With Plate I and six Figures in the Text. 
I N 1830 Nees von Esenbeck described, under the name Monoclea 
Blumei , a remarkable Liverwort collected by Blume in Java. Later 
the plant was removed from the genus Monoclea and named Calobryum. 
For nearly sixty years no further collections of the plant were made, 
when it was rediscovered in Java by Goebel. 1 More recently it has been 
collected by several botanists in the same location, and is also reported 
from Sumatra. 2 
In 1906 the writer spent several months in Western Java, and near 
Tjibodas, on the slope of Mount Gedeh, made several collections of this 
plant. Material from Mount Salak, near Buitenzorg, where Goebel’s 
specimens were obtained, was also examined. 
In addition to the Javanese species, there are two others known 
at present — C. Mnioides , (Lindb.) St., comes from Japan, and C. andinum , 
(Spruce) St., occurs in some of the West Indies and in South America. 
The evidently related genus, Haplomitrium , comprising a single species, 
II. Hookeri, is an extremely rare Liverwort found in Great Britain and 
at a few points on the continent of Europe. These are the only members 
of the family Calobryaceae, whose relationships with other Hepaticae 
are very obscure. 
Goebel 3 has given an excellent account of the more important 
vegetative structures of Calobryum , but details are lacking of the develop- 
ment of the reproductive organs and embryo ; and as the writer’s collection 
afforded very satisfactory material of both male and female plants, as well 
as a small number of embryos and young sporophytes, it seemed worth 
while to investigate these somewhat carefully for comparison with the other 
Hepaticae. 
1 Goebel, K. : Morphologische und biologische Studien. Ann. du Jardin Botanique de Buiten- 
zorg, ix, 1891. 
2 Schiffner, V. : Die Hepaticae der Flora von Buitenzorg, 1900. 
3 Loc. cit. 
[Annals of Botany, Vol. XXXIV. No. CXXXIII, January, 1920.] 
B 
