Calohryum Blumei , N. ab E. 9 
a definite wall from the lower part, forming the ventral canal-cell. In 
several cases noted, both the ventral canal-cell and the cell enclosing the 
egg showed a very conspicuous thickened membrane (Text-fig. 5, D), which 
would seem to preclude any possibility of fertilization. In other cases the 
egg degenerated, and there was some evidence that the ventral canal-cell 
might be fertilized instead of the egg. 
The great similarity in the development of the archegonium and 
antheridium is of interest in connexion with the question of the homologies 
Text-fig. 5. A, Surface view of the neck of an older archegonium, showing the torsion of the 
neck-cells ( x 280) ; b, cross-section of the archegonium neck, showing four peripheral cells ( x 280) ; 
C, venter of a ripe archegonium ; D, venter of an archegonium in which the ventral canal-cell and 
the egg-cell are enclosed in a thick membrane ; E, apex of the neck, seen from the surface ; f, three 
cross-sections of a young archegonium — 1, 2, near the base : 3, the neck ; G, an archegonium, with 
four neck canal-cells, in which no cap-cell had been formed ; H, apex of an abnormal archegonium, 
suggesting an antheridium. 
of these organs. Goebel 1 has argued that the two are homologous, and 
this view would certainly be strengthened by the marked resemblances 
between the archegonium and antheridium in Calobryum. 
The Embryo. 
The material collected by the writer showed only a very few young 
embryos, so that it was not possible to determine with certainty the 
succession of divisions in the earlier stages. 
The youngest specimen found is shown in Text-fig. 6 , A. It corre- 
sponds pretty closely with the usual type of the Jungermanniales, i. e. there 
are apparently two transverse divisions before any longitudinal walls are 
formed. The basal cell, as is usual in the Jungermanniales, probably takes 
1 Goebel, K. : Uber Homologien in der Entwickelung mannlicher und weiblicher Geschlechts- 
organe. Flora, xc. 279-305, 1902. 
