1 12 Salisbury . — Variation in Anemone apennina, Z., and Clematis 
homogeneous in thi^ respect. The data obtained from several hundred 
inflorescences were compared from this point of view, and the marked 
differences which those from the same branch may exhibit, both as 
regards variation range and the position of the mode, is illustrated by 
the curves for four separate inflorescences of the same plant reproduced 
in Fig. 6 . 
The marked trimery exhibited by the gynaeceum of this species is 
all the more interesting since the genus is characterized by a relatively 
primitive carpel in the sense that the functionless rudimentary ovules are 
retained (cf. Warming, 1913). One may hazard the suggestion that the 
marked trimery is associated with this gynaeceal conservatism. In respect 
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Fig. 8. 
to this trimery the individual inflorescence bears much the same relation to 
the plant as a whole as does the * pure line ’ to the homozygous strain 
from which it is selected. This tendency towards a segregation within the 
soma of the individual may be responsible for the fact that if one selects seed 
from flowers with a high number of perianth segments the offspring exhibits 
a higher average than do those raised from seeds produced in flowers of the 
same plant with a low perianth number. In Fig. 8 is shown a diagrammatic 
representation of a branch bearing a number of small inflorescences, and by 
its sides the variation curve for the gynaecea. It will be apparent that the 
range is relatively narrow, and in the different flowers of the individual 
inflorescence the total number of carpels exhibit a striking similarity. 
Here too, as indeed is generally the case, the terminal flower of a dichasium 
usually contains a greater number of carpels than the lateral flowers 
corresponding to it. Their relation is probably largely dependent on 
conditions of nutrition, the first-formed flower being naturally favoured 
in this respect. Exceptions are nevertheless by no means infrequent, and, 
especially in crowded inflorescences, the gynaeceum of the peripheral lateral 
