50 
Sinnott.— The Morphology of the 
ovule, which is eventually forced up into a partially erect position 
(Diagram 5, A). At maturity the bract is difficult to make out, and 
the epimatium appears as merely a thin sheath around the lower third 
of the ovule. The vascular system of the mature cone, investigated only 
in D. cupressimim , is at its base very similar to that of D. Bidwillii 
(Diagram 6, b). The base of the ovule has been left very near the apex of 
the cone, and the two bundles to the ovule, which fuse and become in- 
versely oriented as in other cases, pass directly into the chalaza after a very 
short course in the base of the epimatium (Diagram 6 , c and d). The 
integument is entirely free from the nucellus (Diagram 6 , e), and since it is 
now the only protective coat of the ovule, it is much firmer than in the 
other members of the Podocarpineae which we have examined, and 
has become differentiated into two layers, a narrow but very firm outer one 
of heavy-walled stone cells, and a less rigid, thinner-walled one within. 
These seem to correspond with the hard and soft layers in the ripe integu- 
ment of Eupodocarpus. 
The resemblance in structure and orientation to the seed of Podocarpus 
which is displayed by the young ovules of this section of Dacrydium 
suggests that we have here to do with an instance of recapitulation, and that 
the erect and naked condition of the ovule at maturity is not primitive, but 
has been recently acquired. 
This description of the anatomical features of the female strobilus of 
Podocarpus and Dacrydium agrees in most particulars with the observations 
of Van Tieghem, Strasburger, Stiles, Miss Gibbs, and others. The marked 
structural resemblance between the cone of these genera and that of the 
Abietineae is noteworthy and will be discussed more fully under our 
consideration of the relationships of the Podocarpineae. 
Saxegothea and Microcachrys were not investigated by the writer, but 
good accounts of the strobilar anatomy of the two genera have been 
published by Thomson (17), Tison(18), and Stiles (11). 
The female cone in Saxegothea consists of a considerable number 
of spirally arranged fertile bracts above and of numerous sterile ones below, 
which pass gradually into the vegetative leaves. The whole structure is 
thus more cone-like in appearance than anything we have yet described in 
the family. Each fertile scale bears near its base a single ovule which is 
semi-erect from the first and is provided with a thin epimatium entirely 
separate from the integument. Tison calls attention to the dissimilarity 
in structure between the tissue just below the insertion of the ovule (and of 
which the epimatium is obviously a continuation) and that of the rest of the 
bract. From the vascular supply of the cone axis, which is formed by a ring 
of bundles and is surrounded by a row of cortical canals, a single vascular 
strand passes off to each bract whether sterile or fertile (Diagram 8,e). In 
the case of the latter, however, two bundles, one from each end of the foliar 
