94 Woodburn . — Spermatogenesis in Blasia pusilla , L. 
this granule is the result of a concentration or differentiation of cytoplasmic 
material. No evidence is at hand to show its identity with a previously 
formed body, or that it is of nuclear origin. 
Quite recently Wilson (’ll) has published the results of a study of 
Mnium and Atrichum , two members of the Musci, and Pellia , one of the 
Hepaticae. The latter is of special interest in this connexion, because of 
its close relation systematically to Blasia. Both belong to the anacrogenous 
group of the Jungermanniales, thus occupying an intermediate position 
between Porella of the acrogenous division and Marchantia and Fegatella 
of the Marchantiales. In Mnium and Atrichum he describes the nuclear 
origin of the blepharoplast in the spermatid, the nuclear origin of an 
accessory body, and the same origin for a very unique structure which he 
terms a ‘ limosphere The latter is a mass of chromatic material built up 
from rod-shaped bodies, which have passed out into the cytoplasm from the 
nucleolus. The ‘ limosphere ’ is also present in the spermatid of Pellia . 
No such structure has been discovered in the closely related Blasia and 
Porella , or in Marchantia and Fegetella of the Marchantiales. In other 
respects a number of Wilson’s figures show a striking similarity to corre- 
sponding ones of the writer’s figures of Blasia. In the two Mosses in- 
vestigated, Wilson finds that the blepharoplast may originate from a probable 
centrosome, a conspicuous centrosphere being present during certain stages 
of the ultimate division of the spermogenous tissue. He finds also in all 
three plants a thread distinct in structure from the blepharoplast connecting 
the latter with the nucleus. The author has observed in Polytrichum 
appearances somewhat similar to stages of the ‘ limosphere’ described by 
Wilson for Mnium and Atrichum , but is compelled to interpret them quite 
differently. Escaped sperms of Funaria killed and stained on the slide 
show practically the same form and structure as do those of the Hepaticae 
described by the writer. Figures and discussion of Funaria and other 
Musci will be presented in a later paper. 
Allen (T2) has made a careful investigation of the spermogenous tissue 
of Polytrichum juniperinum , including the last division and the resulting pairs 
of spermatids. These he prefers to term ‘androcytes ’, and the earlier cells 
of the spermogenous tissue f androgones * ; hence the ultimate androgones 
function as androcyte mother-cells. The androgones are characterized by 
the presence of polar plates of kinoplasmic material, or in later cell 
generations by groups of ‘ kinetosomes ’ instead of plates. In either case, 
previous to cell-division, a single plate or group of kinetosomes divides, 
forming two, each taking up its position respectively at the pole of the 
succeeding spindle. In the androcyte mother-cell, instead of a plate or 
a group of kinetosomes, a single ‘ central body ’ of quite similar behaviour 
occurs. Surrounding the central body are radiating fibres, some of which 
extend away from the nucleus towards the periphery of the cell. Connected 
