506 Agnes Arber. — On the Structure of the Androecium in 
that each terminal gland in the staminode of Parnassia palustris is equivalent 
to an anther. Drude has indicated a possible process whereby a staminode, 
such as that of Parnassia palustris , might be imagined to arise from a 
stamen-phalange such as that of a Hypericum . He points out that, in 
certain species of St. John’s Wort, a large gland occupies the upper part of 
XX E. F. 
Text-fig. 4. a. Transverse section of the upper part of the anther of Hypericum sp. (? olympi- 
cum ) to show a gland (g) occupying the greater part of the connective ( x 34). b-f. Series of transverse 
sections through the essential organs of a flower of Hypericum Elodes , Huds. ( x 34) ; j = style ; 
a. ~ anther ; f. = filament ; c.f. = common filament ; std. = staminode ; 0. = ovary. Owing to the 
oblique position of the lateral stamens of each phalange, their anthers are not completely seen in any 
one transverse section. 
the connective in each anther (Text-fig. 4, a) of the stamen-fascicle. If 
these stamens became abortive by the loss of their pollen-sacs, we should 
be left with a bundle of filaments each terminating in a gland. Drude has 
also called attention to another curious analogy between the androecia of 
Hypericum and Parnassia palustris , namely, that the stamen-fascicles and 
the staminodes respectively tend to persist during the ripening of the fruit. 
