A sco carp i?i Lachnea ere tea. 561 
erythrostoma was meiotic or brachymeiotic, and Harper (Publ. Carnegie 
Inst, of Washington, No. 37, pp. 82-4) states clearly that he regards 
the divisions in the ascus of Phyllactinea corylea as bringing about two 
reductions. 
The study of the first and third divisions, and especially of the third 
telophase , in forms with large nuclei, is essential for the satisfactory determi- 
nation of this question. 
Summary. 
1. The archicarp of Lachnea crete a arises as a branch from one of 
the main hyphae and forms a coil of two or three turns. 
2. From the hypha bearing the archicarp and from neighbouring 
hyphae, branches grow out to form a sheath. 
3. The archicarp becomes differentiated into three regions, the multi- 
cellular stalk, the coiled, multicellular ascogonial portion, and the septate 
trichogyne. The latter elongates considerably, and ultimately branches. 
No antheridium was observed. 
4. The trichogyne becomes emptied, its transverse septa break down, 
and the pores are closed by homogeneous pads. It suggests itself that the 
trichogyne has only recently ceased to function. 
5. The ascogonial region contains numerous nuclei. Very large pores 
are found between its constituent cells, and the nuclei pass freely from one 
cell to another. Ascogenous hyphae arise from the several cells. 
6 . Asci are formed in the usual way. The nuclei show about eight 
chromosomes in the first division. 
7. The nuclei were too small to allow a satisfactory study of their 
behaviour either in the ascogonium or in the later divisions in the ascus, the 
significance of which is discussed. 
8. In the structure of its archicarp Lachnea crete a differs markedly 
from the other investigated species of Lachnea. It resembles several of the 
Ascobolaceae, and shows some points in common with Aspergillus. 
List of Papers. 
1 . Blackman, V. H., and Fraser, H. C. I. (’06) : On the Sexuality and Development of the 
Ascocarp in Humaria granulata. Proc. Roy. Soc., London, B., lxxvii, p. 354. 
2. and Welsford, E. J. (’ 12 ) : The Development of the Perithecium of 
Polystigma rubrum. Ann. Bot., xxvi, p. 761. 
3. Brooks, F. T. (’10) : The Development of Gnomonia erythrostoma , Pers. The Cherry Leat 
Scorch-disease. Ann. Bot., xxiv, p. 585. 
4 . Brown, W. H. (TO) : The Development of Leotia . Bot. Gaz., 1 , p. 443. 
