596 
Saxton. — Contributions to the 
left of each diagram, and the total number below these. The ventral nucleus 
is also indicated in Diagram 5 . It may be noted that it is usual, but not 
invariable, for the lowest tier to consist of only a single cell. 
Only scattered stages were examined beyond this, the chief point of 
interest being that two or three tiers of cells contribute to the suspensor. 
This has been found to be the case also in TJmja (Land (24)) and Torreya 
(Coulter and Land (11)), and is said (Coulter and Chamberlain (12)) to be 
usual amongst ‘Taxaceae’ (Taxads and Podocarps). 
As recorded by Hansen (19), the number of cotyledons in the embryo 
ranges from three to six. Text-fig. 6 shows a transverse section of an embryo 
with three cotyledons, while Text-fig. 7 is from a drawing of an embryo with 
five. The only other embryo dissected had four cotyledons. It may be worth 
Text-fig. 5. — Series of diagrams to illustrate some of the arrangements of cells met with in walled 
proembryos of Tetraclinis. 
noting that two to five cotyledons are found in Juniperus (Coulter and 
Chamberlain (12)), while among Callitroi'deae, W iddringtonia occasionally 
has three, but usually agrees with the other two genera, which invariably 
possess only two . 1 
There is one peculiarity in the mature seed which has apparently not 
been recorded before, namely, that the testa is soft, thin, and membranous. 
There could be no doubt that the seeds examined were mature. They were 
collected at the same time as the youngest cones of the succeeding crop, 
and the cones in which they were found were wide open and had already 
shed most of their seeds (all, except in a few cases). The embryos of 
Text-figs. 6 and 7 were dissected out from two of these seeds, and the 
remainder (about three or four) were planted but did not germinate. 
Certain other points, best seen in, or confined to, prothalli containing 
proembryos may now be discussed. 
1 Lubbock (30) states, incorrectly, that A ctinostrobus pyramidalis has three cotyledons. No 
single case has been seen by me, out of a large number of seedlings (certainly more than one 
hundred), where more than two cotyledons were present. 
