599 
Life-history of Tetraclinis articnlata. 
W. cupressoides. Among various morphological points indicating a close 
relationship between Juniperus and Tetraclinis are the delayed division of 
the microspore nucleus, the structure of the female cone and ovules, and 
some other resemblances noted in detail above. It is noteworthy also that 
occasional development of lateral archegonia, as described in Tetraclinis , 
has also been described and figured by Noren (37) in Juniperus , although 
of a somewhat different type to that met with in Tetraclinis . 
The early development of the ovule of Tetraclinis is very similar to 
that of W iddringtonia (except for the absence of a depression at the apex 
Text-fig. 9.— Part of a map of the world, showing the distribution and suggested phytogeny of the 
Callitro'ideae and of certain Cupresso'ideae. 
of the nucellus in the latter), while the development of lateral archegonia in 
the prothallus has become a fixed character in W iddringtonia, no apical 
archegonia having been ever observed. In W iddringtonia, however, the 
position and extent of the lateral groups is much more variable than in 
Callitris and Actinostrobus. It may also be noted that the development of 
the proembryo, as far as has been seen, while substantially similar in 
W iddringtonia, Callitris , and Actinostrobus , is less constant in W iddring- 
tonia than in the other two genera, and differs somewhat less widely from 
the Cupresso'ideae type. The variability of the Tetraclinis proembryo has 
already been pointed out. 
Points such as these suggest that the phylogeny of the three genera of 
the Callitro'ideae, and of the two genera of Cupresso'ideae to which they are 
considered to be most nearly related, is somewhat as shown in Text-fig. 9 . 
