200 Wager . — The Life-history and 
has become arc-shaped, and appears of a peculiar spongy texture. The nucleoplasm is in contact 
with it. 
Fig. 29. Young sporangium with two nuclei, each with a sickle-shaped mass of chromatin and 
a smaller granule or group of minute granules. 
Fig- 30. Young sporangium with two nuclei in process of division. Near each nucleus is a 
quantity of chromatin, probably extruded from the nuclei for purposes of metabolism. This is 
probably derived from the large sickle-shaped chromatin mass or nucleolus which is found in the 
resting stage. 
Fig. 31. Sporangia with four nuclei, three of which are shown. They appear to have just 
finished dividing. A few fine fibrils are seen extending between the pair of nuclei shown, and the 
groups of chromosomes in the newly constituted daughter nuclei are visible. There are two deeply 
stained chromatin masses in the cytoplasm. 
Fig. 32. Sporangium with four nuclei, each with peripheral chromatin mass and nucleoplasm. 
Fig. 33. Sporangium with four nuclei, all in process of division. Equatorial plate just forming. 
The lateral walls of each nucleus prominent and deeply stained. At the poles of the nuclei are well- 
marked radiating striae. 
Fig. 34. Upper two-thirds of a sporangium with four nuclei, all of which are shown in the part 
drawn. Two large, deeply stained granules and two smaller ones, probably extruded from the nuclei 
during division. 
Fig. 35- Sporangium showing the reconstruction of the daughter nuclei. A few delicate strands, 
the remnants of the spindle, are still visible, extending between the young nuclei, and near these are 
large stained granules, probably the remains of the nucleoli. 
Fig. 36. Sporangium with eight nuclei. The nucleolar body forms a sickle-shaped mass at the 
periphery of each nucleus. 
Fig. 37. Portion of sporangium with 32 nuclei. A deeply stained arc of substance is applied 
against the nuclear membrane of each. The granular or thread-like substance in the centre is 
probably just about to break up into chromosomes. 
Fig. 38. One half of a sporangium with 32 nuclei, all in process of division. The spindle is 
clearly shown, and at the poles of some of the nuclei kinoplasmic masses (centrosomes ?) with 
radiating striae are to be seen. The nuclear membrane is thickened laterally. One of the nuclei 
shows clearly ten chromosomes. 
Fig. 39. Sporangium with 32 nuclei, all dividing. Apparently a later stage than Fig. 38. The 
nuclei are flattened slightly at the poles, where the kinoplasm is prominent with radiating striae. No 
granules which could be distinctly called centrosomes were visible in the kinoplasmic masses, although 
these sometimes appeared to be granular. 
Fig. 40. Young sporangium with eight nuclei, all in division stage, showing equatorial plate, 
spindle, and appearance of radiating striae at the poles of the nuclei. The spindle seems to be 
formed out of the lightly stained fine network in the nucleus, and appears before the wall breaks 
down at the poles. 
Fig. 41. Portion of sporangium containing about 32 nuclei in state of division. The drawing 
shows an optical section of a portion with eight nuclei. Four of the nuclei, being at a different 
level, are not shown. Between the groups of daughter chromosomes in each case is an oval or 
spherical mass slightly less deeply stained than the chromosomes, probably the remains of the 
nucleolus. 
PLATE XVIII. 
Fig. 42. Portion of a sporangium in which there were probably 64 nuclei, all in a late stage of 
division. The spindle fibres connecting the groups of daughter chromosomes are visible, and near 
each is a deeply stained granule, probably extruded from the nucleus during division. 
Fig. 43. Portion of sporangium with 16 nuclei just divided. Eleven of these are shown in the 
drawing. Between the pairs of nuclei delicate fibres are still visible, and the cytoplasm contains 
some deeply stained granules. 
Fig. 44. Portion of sporangium just before the segregation of the zoospores. There were 32 
nuclei present, 16 of which are shown. 
