480 Hill . — TVz^ Floral Morphology of the Genus Sebaea . 
received from Dr. Marloth, which he had read at a meeting of the Royal 
Society of South Africa on September 1 6, 1908. In this paper he described 
the secondary stigma in Sebaea exacoides. 
On receiving Dr. Marloth’s full paper, 1 published in July, 1909, I found 
that he had reached the same conclusion, with regard to the stigmas of 
Sebaea exacoides , to which I had been led from an examination of the 
whole genus and by the series of experiments shortly to be described. 
Although the results of this work have to some extent been forestalled 
by the publication of Dr. Marloth’s interesting observations made in South 
Africa, they seem worthy of being placed on record as an independent 
account of the peculiar floral structure which obtains in the genus. As 
a further extenuating circumstance, it maybe pointed out that the investiga- 
tion has been carried out on lines different from those followed by 
Dr. Marloth. 
The Natural Order Gentianaceae shows several peculiarities of stigmatic 
arrangement in the different genera, which appear to be correlated with the 
structure of the corolla. Examples of dimorphic heterostylism are found 
in the genera Exochaenium, 2 Hockinia , and to a slight extent in Lisianthus , 
all genera in which there is a well-marked corolla tube, and it seems possible 
that this condition may also occur in the genus Sebaea itself. Then in 
Pleurogyne , 3 with its widely open and almost polypetalous flowers, the 
stigmatic surfaces form two bands down the sides of the ovary and the 
normal apical stigma appears to be more or less functionless (see Figs. 
1 and a, PL XXXV). 
In Sebaea , where there is a distinct and more or less cylindrical 
corolla tube, the style is usually elongated. The stigma at its apex may be 
clavate, capitate, or in many cases shortly bilabiate. Except in the two 
species included in the sub-group Brevistylae, 4 .S', spathulata (PL XXXV, 
from Mr. Burtt Davy, and some collected from the dried material sent over from the Transvaal 
Herbarium for identification. From the seeds so obtained numerous plants were raised and afforded 
ample material for experiment. 
On October 7 of the same year Dr. Marloth kindly sent seeds of Sebaea ( Belmontia ) cordata , 
and wrote as follows : ‘ The box also contains some seeds of Belmontia cordata, as I heard from 
Dr. Bolus that in connexion with the Part on Gentianaceae for the “ Flora Capensis ” you wished to 
raise some plants from seed. I take the liberty of enclosing an abstract of a paper on some bio- 
logical features of Belmontia .’ 
To complete the record of the seeds sown at Kew a packet of seed of Sebaea ambigua was 
received from Mr. Worsdell, from the Cape, in April, 1910, which also yielded some interesting 
seedlings, and I desire here to acknowledge my indebtedness to all who have kindly assisted me by 
sending seeds from South Africa. 
1 Marloth, R. : ‘ A Diplostigmatic Plant,’ Sebaea exacoides , (L.) Schinz, in Trans. Roy. Soc. 
of S. Africa, vol. i, pt. 1. 
2 See A. W. Hill in Kew Bull., 1908, pp. 336-40 with Plate. 
3 See Engler und Prantl, iv. 2, p. 87, Figs. D, E, esp. P. carinthiaca. Huxley, in his paper 
on the Gentians in Journ. Linn. Soc., xxiv, pp. 103, &c., refers to the types of corolla, but does not 
describe the stigmatic arrangements in any of the genera. 
4 A. W. Hill in Kew Bull., 1908, p. 320; FI. Cap. iv. 1, pp. 1062, 1092. See also p. 487. 
