Hill.— The Floral Morphology of the Genus Sebaea. 483 
it can be seen that the stigmatic tissue is continued down the style from 
the point of union of the edges of the stigmatic lobes. These two decurrent 
lines or bands of papillae on opposite sides of the style are thus at right 
angles to the surfaces of the stigmatic lobes, and undoubtedly represent the 
first stage in the evolution of the definitely separated secondary stigmas. 
In this species the apical stigma scarcely projects beyond the anthers, and 
the decurrent papillate bands merely extend the stigmatic surface to a 
point slightly below the base of the anthers, being continuous above with 
the edges of the lobes of the stigma. 6\ ovata, R. Br. (Benth., FI. Austral., iv, 
Text-fig. i. Diagrammatic figures to show the supposed mode of origin of the paired lateral 
stigmatic patches in Sebaea. i. A simple bilabiate stigma, the inner surfaces of the lobes (s') being 
stigmatic. 2. As Fig. t, showing the edges of lobes (se) recurved and stigmatic. 3. The recurved 
edges of the terminal stigmatic lobes are continued down the style (Is.) as lateral stigmatic areas. 
4. The lateral stigmatic patches, consisting of two ridges, have become separated from the terminal 
stigma. 5. A stigma tending to become clavate by the fusion of the lobes ; the edges are recurved 
and continued down the style as decurrent stigmatic bands. 6. A clavate stigma, the lobes being 
folded together, the edges forming the stigmatic band over the top of the style and continued below as 
the lateral or secondary stigmatic patches (is.). 7. A more typically clavate stigma, the lateral 
patches slightly detached from the terminal stigma. 8. The lateral secondary stigmas ( Is .) widely 
separated from the terminal stigma. 
p. 371), shows a similar arrangement to S. khasiana, and, although the 
style in this species overtops the anthers, the stigmatic surface is decurrent 
from the edges of the lobes to the level of the base of the anthers, and the 
secondary stigmas take the form of twin ridges bearing papillae on each 
side of the style (PI. XXXV, Fig. 3). Sebaea membranacea (Figs. 22 and 23) 
and .S', ecarinata 1 (Figs. 19 and 20), two species found by Professor H. H. W. 
Pearson during the Percy Sladen Expedition to the Orange River, show 
the secondary stigmas in close juxtaposition to the apical one, but definitely 
separated from it as distinct organs. The separation of these patches 
of stigmatic papillae may thus be regarded as having been effected by the 
1 Gentianaceae in Annals of the S. Afr. Mus., ix, pt. 1 1, pp. 57, 58. The lateral stigmas in 
S. acutiloba , S. Zeyheri, S. micrantha, and S. intermedia are also nearly confluent with the terminal 
stigma. See FI. Cap., iv. 1. 
L 1 
