Anatomy of Pherosphaera Fitzgeraldi , F. v. M. 313 
The green tissue immediately within the hypoderma deviates from 
typical palisade-parenchyma as regards both shape and orientation of the 
cells and dimensions of the intercellular spaces. The general design of this 
loose tissue is that of two layers of elongated cells, simulating palisade-cells, 
and stretching between the hypoderma and the central strand of the leaf. 
These cells, and particularly those forming the outer layer, are mainly 
perpendicular to the leaf-surface, but those radiating from the central strand 
towards the lateral angles (in transverse section) deviate from this design so 
greatly that in the angles they are actually tangential to the local curved 
surface. The cells forming the outer green layer frequently are lobed either 
Transverse section of leaf of Pherosphaera Fitzgeraldi (slightly diagrammatic). 
laterally or basally, or have dilated bases ; this is especially the case within 
the upper face of the leaf, so that here in transverse section a number 
of structures may be seen that apparently are short cells but really are the 
basal portions of long cells. The green cells of the outer series attain their 
maximum length towards the upper face of the leaf. 
The cells forming the inner layer of palisade-like tissue on reaching the 
central strand may bend so that their inner portions run parallel to the 
length of the leaf and thus contribute to the outermost layer of the central 
strand : such cells consequently lose their palisade-like form, and in 
transverse section are liable to be mistaken for tissue confined to the central 
strand. 
The cells forming the outermost layer of the central strand contain 
chlorophyll, are elongated in the direction of the leaf-axis, and may emit 
