536 DutL — Pityostrobus macrocephalus , L. and H. 
3. The O villiferous Scales . 
The tissue of an ovuliferous scale consists chiefly of rather large, very 
thick-walled cells, often showing contents. In the surface layers the cells 
are smaller and more strongly sclerized. It is noticeable, however, that the 
scale as a whole, and particularly the basal region, as seen in transverse 
section, does not show such a large development of sclerotic tissue as seems 
to be usual in recent cones of a similar size. The difference in the thicken- 
ing layers of the upper and lower surface is most clearly exhibited in the 
sections from the Sedgwick Museum cone. The lower layers of the scale 
consist of very narrow elongated sclereids appearing as fine wavy lines in 
a longitudinal section, and as crowded, circular areas with minute lumina in 
a transverse section of the scale. The layers of the upper surface consist of 
larger sclereids with polygonal outlines and frequently contain massive cells 
Text-fig. 2. P. ovatus. Transverse section of ovuliferous scale about half-way up, showing 
disposition and orientation of vascular bundles and chief resin-canals. v.b. t vascular bundle ; 
xyl ., xylem ; phi., phloem ; r.c., resin-canal. Camera lucida drawing. x 6. Cowderoy Col- 
lection, V 8309. 
with peculiar irregularly elongated shapes, which may be compared to the 
idioblasts found in the cone-scales of the Araucarineae, &c. In the larger 
middle region the cells are only moderately thickened, and the whole tissue 
is permeated with a profusion of secretory canals. In P. ovatus the upper 
sclerized region is not so large, nor so clearly delimited from the middle 
region. 
The seed-scale is supplied by two separate vascular strands arising 
independently from the sides of the gap in the main stele, just as Miss Aase 
(1) has recently shown to be the case among many recent conifers. Most 
transverse sections of the cone show the two bundles very clearly (Fig. 6, 
PI. XV). The scale traces quickly broaden out and divide so as to form 
a single series of bundles with phloem uppermost. Text-fig. 2 shows the 
distribution of the bundles in a cross-section about half-way up the vertical 
part of the scale. The small size of the bundles and their frequent arrange- 
ment in pairs are noticeable features. In P. macrocephalus the bundles, 
though more numerous, seem to be even more minute than represented in 
