Davey.— Seedling Anatomy of certain Amentiferae . 585 
Juglans Sieboldiana. This seedling agrees with the preceding species 
in the structure of its cotyledon strands and in their arrangement at the 
cotyledonary node. The traces of the first and second plumular leaves 
in the intercotyledonary plane show double structure with isolated median 
protoxylem, but this last feature is not continued into the epicotyl, where the 
leaf-trace consists of two widely separated endarch bundles. The structure 
of the hypocotyl and of the root is tetrarch and resembles that of the last 
species, but some of the xylem of the lateral cotyledonary strands diverges 
towards the intercotyledonary plane, where it becomes connected with the 
plumular strands. The greater part of the metaxylem of the intercotyle- 
donary root poles is continuous with that of the cotyledonary strands, 
but the central portions are continued up into the epicotyl as the traces 
Fig. 8. Juglans Sieboldiana. Outlines of vascular system. A. At cotyledonary node. B. At 
a lower level in the hypocotyl. cot. = cotyledonary traces. Arrows indicate cotyledonary and 
intercotyledonary planes. 
of the first two leaves. In other respects, such as the presence of a large 
pith and early secondary thickening, and also in the fact that metaxylem 
from later plumular leaves may persist in the diagonal planes, the seedling 
is similar to that of Juglans nigra. 
Juglans cinerea. The cotyledon petioles show the usual five strands, 
but the central one is extremely small, and the constituents of the 
laterals are not compactly united. These enter the axis as groups of two 
or three approximated bundles at the diagonal corners. The plumular 
strands are very slightly differentiated, but show doubleness of their phloem. 
In the hypocotyl, the cotyledonary poles are supplied by the central strands 
together with adjacent parts of the lateral groups, the remaining portions of 
which are connected with the intercotyledonary poles as in the normal 
tetrarch type. No plumular tissue takes part in the primary organization of 
root poles. 
Juglans regia , This seedling showed tetrarchy in root and hypocotyl 
