SKULL OF THE PLEISTOCENE TAPIR. 
67 
RELATIONSHIP TO EXISTING SPECIES. 
While the small tapir of the Ashley River deposits may thus 
provisionally be united with that from Florida, it is certain that the 
Florida tapir cannot be united with the. recent Tapirus terrestris of 
South America. The skull differences have already been indicated. 
The face of Tapirus terrestris is relatively elongated; the crest is 
strongly developed, especially in old and mature individuals. The 
spiral groove of T. terrestris is more deeply intrenched into the 
frontals than is that of T. veroensis. The lachrymal pit of T. vero- 
ensis is lacking in T. terrestris. 
Owing perhaps to the conservative pattern of tapir teeth, the 
differences in tooth structure between the Recent and Pleistocene 
species are less pronounced than are some of the other characters 
of the skeleton. However, where there are such notable skull dif- 
ferences, some characters of systematic value may be expected in 
the teeth. Such is the case. Notwithstanding the elongation of 
the face, the molars and premolars of T. terrestris are compressed, 
giving relatively increased transverse measurements ; the diastema, 
on the other hand, is relatively increased. Other differences may 
be noted. The cingulum at the front outer side of the cheek teeth 
of 7 . veroensis is stronger than is that of T. terrestris. The first 
upper premolar of T. terrestris is much widened at the posterior 
margin. On the other hand, the anterior division of Pm 2 is not so 
wide as is the corresponding division of the same tooth of T. vero- 
ensis. The third molar of the Florida tapir is larger than that of 
the recent South American species. On both molars and premolars, 
the ridge or crest which runs to the posterior side of the paracone 
meeting the anterior walls of the metacone is much stronger on the 
teeth of the T. veroensis than on those of T. terrestris. The tubercle 
which is found in the Pleistocene tapir at the inner side of the upper 
cheek teeth between the transverse crest, is lacking, or nearly so, in 
the Recent T. terrestris. 
These differences, both in the skull and the teeth, separate this 
fossil tapir from the existing South American species. 
ASSOCIATED FOSSILS. 
The fossils of the marine shell marl include as known at present 
61 species of mollusks, of which one species is apparently an extinct 
