104 Hill.—The Structure and Development of 
Flower-development , &c. 
1. The various parts of the flower are arranged in alternate 
whorls of three, which arise in acropetal succession. The 
anterior calyx-lobe is the first organ to arise. 
The ovule is basilar in origin. 
2. The embryo-sac often contains an increased number of 
antipodal cells, the number varying from three to fourteen, 
as far as has been seen. 
3. The polar nuclei do not appear to fuse till after ferti¬ 
lization has taken place. 
4. The embryology follows a normal course strongly 
resembling that of other Monocotyledons. A basal cell is 
developed, and it appears that this does not undergo division. 
5. The embryo does not bend over, as in Alisma\ the 
stem-apex is developed laterally. 
6. An endosperm is formed, the nuclei of which are not 
separated by means of cell-walls; they are very distinct, 
lining the inner wall of the embryo-sac. 
The greater part of this research was carried out in the 
botanical laboratories at the Royal College of Science, and 
I should like here to express my best thanks to Professor 
Farmer for the help and advice he has given me. 
My thanks are also due to the Director of the Royal 
Gardens, Kew, and to Dr. Scott, Hon. Keeper of the Jodrell 
Laboratory, for allowing me to work in the Jodrell Laboratory 
during part of the summer vacation. 
