i6o 
Notes. 
in the cortex. Lower down, just beyond the transitional region between 
stem and root, and where root-structure already prevails, the central 
stele is of considerable width, and the constituent bundles farther apart, 
owing to the expansion of the parenchymatous tissues. The large 
bundles composing the secondary tissues of the stele are now in places 
connected by delicate, few-layered bands of secondary vascular tissue, 
whose course is rather difficult to follow. 
So far the structure is normal enough. But outside the central 
normal stele are seen, in transverse section, other strands which, 
there can be no doubt, represent the extrafascicular vascular tissue 
of Cycas , Macrozamia, and Fncephalartos. Now this extrafascicular 
vascular tissue, which, at this youthful stage, only occurs outside 
a small part of the circumference of the central stele, has this 
peculiarity: that it consists of two distinct parts, viz. an outermost 
normally orientated and an innermost band whose parts are abnormally, 
i. e. inversely orientated, with the xylem directed outwards and the 
phloem inwards. Of these two well-marked portions of the extra¬ 
fascicular tissue, the inner—or that with inverted orientation—is, at 
this stage of the plant’s life-history, the best developed of the two. 
Both portions, in accordance with the youthful age of the plant, are 
as yet in a very rudimentary stage of development, the xylem and 
phloem being in fact but just sufficiently advanced in differentiation 
for the determination of the above salient characters. 
The above structure I regard as homologous with, although a 
slight modification of, that described by Gregg in the root of Cycas 
Seemanni, Al. Br., and by myself in the root of Cycas revoluta, Thunb., 
and the stem of Macrozamia Fraseri, Miq. 
It appears to me, moreover, to distinctly support the view put 
forward in the paper dealing with the latter plant, that the vascular 
tissues (at least, the extrafascicular zones) are derived by modifi¬ 
cation from those of plants like the Medulloseae, which normally 
possess vascular strands, each of which exhibits the mutually-inverted 
portions above described. 
W. C. WORSDELL, Kew. 
A NEW CARDIOCARPON-BEARING STROBILUS.— The 
writers have latterly each met with specimens of Cardiocarpon, as 
figured by the late Professor Williamson and other authors, but 
attached to a central axis in a strobiloid form (resembling a Lepido- 
