Saccharomyces anomalus {Hansen). 239 
film is formed, which originally consists entirely of oidia, 
but later chlamydospores appear and give to the film a 
yellowish colour. 
On purely morphological grounds it will be noticed that 
there are certain resemblances between the two forms, but 
with reference to the mycelial formations, it would appear 
that here there is a distinct difference. Fig. 12, however, 
taken from the edge of a colony of S. anomalus , shows 
that there is a great tendency in this organism to produce 
a mycelium on a solid and dry medium. The asci in the 
two cases present considerable differences however. In S. 
anomalus, as far as we know, an ordinary vegetative cell 
becomes developed into an ascus under suitable conditions, 
while in E. decipiens the ascus is developed from a side- 
branch of the mycelium, i.e. in a somewhat definite position. 
This distinction, however, must perhaps not be pressed too 
much until definite knowledge is obtainable as to the power 
of every vegetative cell of the Yeast to form ascospores, and 
it is conceivable that it points merely to a slightly more 
specialized condition in the case of E. decipiens. The asco¬ 
spores of both forms resemble one another completely in 
shape. Each ascus in E. decipiens typically produced four 
spores, while four is the most usual number in the form of 
*S\ anomalus which I have described. The germination of 
these spores, however, differs in a marked degree owing to the 
formation of a pro-mycelium by E. decipiens. The question 
may be raised whether this difference is as important as it 
appears at first sight. Hansen ( 1 ) has shown that the spores 
of S. Ludwigii , a species which is nearly related to S. ano¬ 
malus, produce a sort of pro-mycelium on germination, while 
Reess (20) pointed out that the spores of Taphrina Pruni 
(Tub) germinate in the same way as ordinary Saccharomyces 
spores, i.e. by budding. Brefeld ( 18 ) has shown the same 
for Taphrina rhizophora (Johans). The genus Taphrina is 
usually regarded as related to Endomyces. 
From these examples, then, it does not follow that the 
production and non-production of a pro-mycelium are facts 
