288 Lang.—Development of Cycadean Sporangia , 
the majority of the sporogenous cells surrounding the embryo- 
sac simply become disintegrated and absorbed, the outermost 
form a more definite tapetal layer. This tapetum, while 
persisting longer than the more internal cells, ultimately 
disappears. 
The course of the vascular bundles in the ovule is most 
easily followed at this stage, and may be briefly described. 
The general arrangement of the bundles in the sporophyll 
has been described and figured by Worsdell 1 . As he points 
out, the bundles passing to the ovule arise by division of 
a single branch from the vascular system of the lateral part 
of the sporophyll. This branch always divides into two 
bundles, one of which may again divide before reaching the 
base of the ovule. Thus two or three bundles of collateral 
structure enter the ovule. These bundles divide up at the 
base of the ovule into sixteen small bundles, eight of which 
bend out at once to form the vascular system of the outer 
layer of the integument, while the eight bundles alternating 
with them pass into the innermost layer. The bundles of 
both the inner and outer series, which thus take their origin 
at the same level, undergo subdivision though not to any 
great extent; those of the outer layer extend to the micropyle, 
while those of the inner series stop at the separation of 
nucellus from integument. 
The remaining observations on the changes taking place in 
the ovule of Stangeria were made on two cones produced 
in the Edinburgh Botanic Gardens, and pollinated about the 
middle of February 1899. So far as can be judged, the cones 
at the date of pollination must have borne ovules of about 
the age of those last described. Dr. Balfour informs me that 
they were then erect on stiff peduncles, and considerably 
smaller than they became after pollination. The increase in 
size took place shortly after pollination, and was attended by 
a change in the position of the cones, which became pendulous. 
When the plant came under my observation in the beginning 
of July, the cone, excluding the peduncle, measured 14 cm. in 
1 Annals of Botany, vol. xii, p. 215, PI. XVII, Fig. 5. 
