49 8 Shove.—On the Structure of the 
in the leaf-bases and stipules have been examined, but only 
to a small extent, as work in this direction is being done by 
Mr. Brebner, of the University College, Bristol. 
I should like here to express my hearty thanks to 
Mr. Seward, of Emmanuel College, Cambridge, for supplying 
me with the material upon which the following account is 
based, and also for his ready help and kindly interest 
throughout my work. 
Previous Work on the Anatomy of Angiopteris. 
The first important account of work on Angiopteris occurs 
in the ‘ Monographic des Marattiacees,’ by De Vriese and 
Halting 1 . In this paper several facts are stated in detail, but 
there is little investigation into the actual course of the 
vascular strands. The mistake of Harting’s in confusing 
roots traversing the stem with stem-bundles is too well known 
to be more than mentioned. 
In 1863 appeared the important paper by Mettenius, ‘ Ueber 
den Bau von Angiopteris.’ Mettenius investigated a stem of 
Angiopteris evecta , Hoffm., but unfortunately the stem was 
decayed at the base, and had shown at the apex no signs 
of life for fifteen months 2 , facts which must be taken into 
consideration in comparing his results with those of later 
workers. 
In this paper the course of the bundles is worked out in 
great detail; briefly stated, the vascular tissues are con¬ 
sidered to form funnel-shaped zones with their lower ends 
in the axis of the stem, and their higher parts continued out 
into the leaves. A transverse section of these funnels would 
give rise to the rings of scattered bundles such as are seen 
in actual sections of the stem. 
Segments from the outermost zone pass off into the leaves, 
and the gaps thus made are filled up by corresponding 
segments from the next inner zone. This method of com¬ 
pensation goes on from zone to zone. Mettenius further 
De Vriese and Harting (’ 53 ). 
2 Mettenius (’ 64 ), p. 504. 
