509 
Stem of Angiopteris evecta. 
six strands arranged as in Fig. 9 ; these divide and give off 
branches from their inner surface which turn towards the 
flat side of the leaf, so that at a distance of about 2 cms. 
from the position of the first section, they represent the 
appearance shown in Fig. 12. 
The initial strands are in general of the same size; the two 
large side strands described and drawn by Mettenius 1 do 
not appear to be present. 
The series of transverse sections shown in Figs. 9-12 corre¬ 
sponds to leaf I in Fig. 6 ; this particular leaf was one of 
the largest on the stem. 
In Figs. 7 and 8 is shown the actual network in the leaves 
formed by the branching strands ; Fig. 7 shows the vascular 
tissue of a leaf from the dorsal, Fig. 8 those of a leaf from 
the ventral side of the stem; in both cases the strands had 
not attained their full development. The cut ends of the 
steles lie in the same plane, being exposed in their present 
position when the corresponding leaf-base was cut from the 
stem ; in Fig. 7, however, the strands A and B belong to 
other leaf-bases. Fig. 8 shows a root (r) originating at the 
base of a leaf-trace bundle. The different character of the 
meshes of the network in the leaf according to its position 
on the stem is clearly seen in the drawings. 
It is interesting to compare the course of the vascular strands 
in A. evecta with that in Kaulfussia and Marattia described 
by Kiihn 2 . 
Kaulfussia possesses a single-meshed cylinder of vascular 
tissue enclosing a central strand ; from the outer zone 
segments are given off to the leaves, and compensation for 
these is afforded by segments from the central strand. 
The vascular tissue in Marattia is more complicated; here 
there is a central strand enclosed by two meshed zones, from 
the outermost of which segments are given off to the leaves, 
while a compensating segment from the second zone fills 
up the gap in the first, and a branch from the central strand 
fills up the gap in the second. Angiopteris differs from 
1 Mettenius (’64), p, 512 . 2 Kiihn (’89). 
