66 Bower.—Studies in the Phytogeny of the Filicales . VII. 
Gymnogramminae, Cheilanthinae, and Adiantinae. 1 But it must be asked, 
Are all of these Ferns to be traced phyletically to a two-lipped sorus, as in 
the case of the Pteridinae, or as I prefer to call them, the Pterid series ? 
It is only on the basis of detailed examination and comparison that this 
question can be resolved. But such examination must be held over 
for the present, with the simple remark that there is no apparent reason 
for holding as justifiable the assumption that they have been so derived. 
Summary. 
1. The stelar ontogeny of Schizaea and Anemia starts from proto- 
stely, a condition retained in the adult Lygodium . It then progresses to 
the medullated monostele, a state retained by the adult in .S'. pusilla and 
rupestris , and apparently also in S. fistulosa. In 6*. dichotoma , and less 
perfectly in S.malaccana, it progresses further to ‘ ectophloic siphonostely \ 
In A?iemia ( §Anemiorrhiza) the development proceeds to ‘amphiphloic 
siphonostely 5 (= solenostely), and in Eu-anemia and Mohria to dictyostely. 
These states may be held as indicating successive steps of stelar advance, 
the ontogeny being less completely carried out in some forms than in 
others. 
2. In Schizaea rupestris and digitata the sporangia have been shown 
to be of marginal origin, and the later-formed indusium originates super¬ 
ficially below the sporangia. This condition prefigures that seen in other 
Marginales. 
3. Within the Pterideae of Prantl there are two probably distinct lines 
of phylesis. The first may be styled ‘ Pterideae bi-indusiatae ’, or better, 
the Pterid series; the second the ‘Pterideae uni-indusiatae’, or better, 
the Cheilanthoid series. The latter will be held over for the present. 
The former, which include Lindsay a, Paesia , P teridium, Lonchitis , Histio- 
p ter is, Anopteris , P ter is , and Acrostichum , L., are traceable from a two¬ 
lipped Dicksonioid origin. Perhaps Saccoloma may also be included. 
4. The Lindsaya-ty^o. of stele has been verified for eighteen species of 
the genus, and may be accepted as a good generic character. The genus 
is further characterized by its ‘ fusion-sorus \ with vascular commissures. 
5. In its ontogeny the fusion-sorus of Lindsay a is actually marginal, 
the indusial flaps originating superficially below it. The sporangia show 
at first a gradate sequence, but soon younger sporangia are interpolated, 
and a mixed condition is assumed. 
6. In Dictyoxiphium the fusion-sorus differs from that of Lindsaya 
chiefly in the absence of the upper (adaxial) indusium. 
7. Except for amendments of detail the stelar system of Saccoloma has 
been shown to accord with Mettenius’s description. 
8. The fusion-sorus of Pteridium is marginal in origin, with two 
1 Compare Diels, Natiirl. Ptlanzenfam., i. 4, p. 254. 
