Campbell—Studies on some East Indian Hepaticae . 333 
Rhizoids of three kinds are present—large smooth-walled ones, some¬ 
what smaller ones with occasional tuberculate thickenings, and much smaller 
ones with numerous tubercles (Text-fig. 9, B). 
The most striking character of D. calcicola is the arrangement of the 
receptacles. As has been stated already, male and female receptacles occur 
upon the same individual, and structurally these do not differ from those of 
other species of Dumortiera. Being of the composite type, the formation 
of a receptacle necessarily stops the growth of the shoot of which it is a part, 
and in most species the receptacle is obviously terminal, and only one 
receptacle is borne on the branch. In D. calcicola , however, as we have 
seen, there are apparently several receptacles formed in succession on the 
same branch, but this is not really the case. Soon after the first receptacle 
is formed an adventitious shoot appears close to the apex of the primary 
shoot on its ventral surface, and this shoot soon develops at its apex another 
receptacle ; and the process may be repeated several times, so that there is 
a series of short branches, each bearing a receptacle, and so linked together 
as to give the appearance of a single deeply-lobed thallus bearing a series of 
dorsal receptacles. 
Sometimes two of these adventitious shoots may develop almost 
simultaneously, thus- simulating a dichotomy of the shoot apex (PI. VIII, 
Fig. 3). The last-formed adventitious shoot may develop into an elongated 
sterile branch. 
As a rule the primary receptacle bears antheridia, while the later ones 
are archegonial; but secondary antheridiak receptacles are not uncommon, 
and not infrequently androgynous receptacles occur. 
The antheridial receptacle (PI. VIII, Figs. 6, 7) is usually 5-6 lobed, 
sometimes quite distinctly triangular in outline (PI. VIII, Fig. 1, B), while 
the female receptacle is nearly circular. Both have the margin beset with 
stiff hairs. On the ventral surface are irregular scales and numerous slender 
tuberculate rhizoids. The scales are especially noticeable in the younger 
stages, the subsequent dorsal growth of the receptacle pushing them under 
so that they are not visible from above. 
The male receptacle and antheridia resemble those of the other species, 
except that they are decidedly smaller and the receptacle is often distinctly 
lobed. The antheridium has the elongated pedicel observed in the other 
species. 
The young archegonial receptacle is somewhat intermediate in form 
between that of D. trichocephala and D. velutina. It is less convex than in 
the former, but more so than in D. velutina. The archegonia have the 
strongly curved neck observed in D. trichocephala (PI. VIII, I^igs. 8, 9). 
All of the female receptacles were sessile; but whether this is the case 
where the archegonia have been fertilized, or whether in such case a pedicel 
would develop as in all other species, is impossible at present to say, since 
