579 
Erythrococca and Micrococca. 
these species, C. fallax , Muell. arg., from Fiji, has been referred to Gymno- 
claoxylon because Mueller could not find glands on the male receptacle. 
The male flowers of the original specimens (Seemann, 394) have, however, 
just as many glands,hirsute at the apex with viscid hairs, 1 as there are stamens. 
In the other species, C. sandwicense, Muell. arg. (Linnaea, xxxiv. 165), there 
are no glands with viscid apical hairs. But associated with the fully 
developed stamens are many smaller and imperfect ones. These imperfect 
stamens show every transition from a filament bearing two free or nearly 
free empty anther cells, to a glabrous receptacular gland. The receptacle 
in C. sandwicense does possess glands, and the circumstance that these glands 
are glabrous does not exclude the species from the section Euclaoxylon. 
That section contains other species with glabrous glands, among them being 
C. parviflorum, A. Juss. (Tent. Gen. Euphorb., 43), the plant on which the 
genus Claoxylon was based. While, therefore, the section Discoclaoxylon , 
Muell. arg., stands, the section Gymnoclaoxylon , Muell. arg., 2 is an integral 
part of Euclaoxylon. 
The section Euclaoxylon, Muell. arg., to which, in 1866, thirty species 
were referred, but which is now believed to include over forty, has been 
found since then to be susceptible of some readjustment. Hooker f., in 1887, 
when revising the Indian species of Claoxylon (Flor. Brit. Ind., v. 410-14), 
was under the necessity of recognizing two sections within what corresponds 
to Euclaoxylon , and under the further necessity of removing from Euclaoxylon 
another species, C. oligandrum , Muell. arg. (Linnaea, xxxiv. 164), from 
Ceylon. With the necessity for subdividing Euclaoxylon we do not here 
require to deal; this can only be appropriately discussed in connexion with 
a critical revision of the genus Claoxylon as a whole. The transfer of 
C. oligandrum from Euclaoxylon does, however, immediately concern us, 
because of the fact that Hooker has found it most convenient to place this 
species in Micrococca. Following Thwaites and Mueller, Hooker has treated 
Micrococca as a section of Claoxylon ; to the consideration of this section, or 
genus, we may now turn. 
1 In C. fallax these glands are thin and flaccid ; each gland is closely applied to and partially 
embraces the base of the filament to which it corresponds. When a flower is examined under 
a simple microscope the appearance presented is practically that described by Mueller of stamens 
with short silky filaments (DC. Prodr., xv. 2, 780). But if a flower which has been softened in 
water be kept under observation, as it becomes dry the glands are seen to separate from the filaments, 
which are perfectly glabrous and are organically free from the glands. 
2 The recognition of a section limited in accordance with the definition which Mueller has pro¬ 
vided for Gymnoclaoxylon is not, on this account, rendered impossible; there are several species 
which since 1866 have from time to time been referred to Claoxylon , where there are no receptacular 
glands. But even if we assume that the reference of these species to Claoxylon be correct and that 
the recognition of such a section be necessary, that section will not include either of the plants 
on which Gymnoclaoxylon , Muell. arg., was based, and therefore will not be Mueller’s section so 
named. 
