On the Gametophytes and Embryo of Pseudolarix. 
BY 
KIICHI MIYAKE 
AND 
KONO YASUI. 
With Plate XLVIII. 
LTHOUGH the gametophytes, fertilization, and embryogeny of the 
Abietineae may, at present, be said to be the best known among 
the Coniferales, there remain still some genera of which the gametophytic 
structures are entirely untouched by investigators. Among the existing 
nine genera of the Abietineae, the genus Pinus has been most thoroughly 
investigated. We have also more or less complete records of the gameto¬ 
phytes of Picea, Adzes, Tsugct, Psendotsuga, and Larix} But, in the 
remaining three genera— Cedrus , Pseudolazdx, and Keetleria —this phase of 
life-history is entirely unknown. In light of the accumulating evidence 
in support of the view that the Abietineae is a very ancient group of the 
Coniferales, 2 it has been thought very desirable that the gametophytic 
phase of all the genera of the group should thoroughly be worked out. 
The present study has been undertaken with this object in view and to fill 
up one of the gaps in our knowledge of the life-history of the Abietineae. 
Psendolarix Kaempferi , the only species of the genus, is a native of 
China and seems to thrive quite well in Southern Europe. The material 
for the present study has been secured from Northern Italy, through the 
kindness of Dr. R. Rovelli in Pallanza, in whose garden a beautiful specimen 
of this tree is growing, and to whom we are very much indebted. During 
the years 1903 and 1904, it was sent from there to Bonn, Germany, where 
one of us (K. M.) was staying at the time. The living male or female cones 
with part of the attached branches were sent two or three times a month 
1 Chief contributions to our knowledge of the gametophytes and embryo of the Abietineae are 
those of Hofmeister (’51, ’58), Strasburger (’69, ’72, ’78, ’79, ’84, ’92), Goroschankin (’80, ’83a, ’83 b), 
Sokolowa (’90), Mottier (’92), Belajeff (’93), Dixon (’94), Coulter (’97), Blackman (’98), Chamber- 
lain (’99), Cavara (’00), Murrill (’00), Arnoldi (’00), Coulter and Chamberlain (’01, ’10), Ferguson 
(’01a, ’01 b, ’04), Miyake (’03a, ’03£), Thomson (’05), Stopes and Fujii (’06), Kildahl (’07), and 
Lawson (’09). 
2 Concerning the anatomical and palaeontological evidences, see Jeffrey (’05, ’08) and Hollick 
and Jeffrey (’09). 
[Annals of Botany, Vol. XXV, No. XCIX, July, 1911.] 
U U 2 
