645 
Ev^bryo of Pseudolarix. 
The female gametophyte is filled with solid tissue early in June, and 
soon the archegonial initials appear at the micropylar end of the prothal¬ 
lium. The archegonia reach the mature size in about three weeks. 
The development and structure of the archegonium do not seem to 
differ much from those of other Abietineae. The number of archegonia 
in a single ovule varies from four to seven, the more usual number being 
five or six. The neck consists usually of eight cells, arranged in two tiers 
of four cells each. 
In the division of the central cell of the archegonium, the chromatic 
contents of the nucleus at first accumulate near the centre of the cavity 
as in synapsis. The ventral canal-cell is formed as a result of the 
division, and it shows immediately signs of disorganization. The egg 
nucleus increases in size soon after its formation, and moves down towards 
the centre of the egg. 
Fertilization takes place about the end of June. The actual fusion of 
the sexual nuclei was not observed. The first two divisions of the fertilized 
egg nucleus take place near the centre of the egg, and the four free nuclei 
of the proembryo move down to the base of the archegonium. The four 
nuclei now divide again, and the walls are then formed between eight free 
nuclei, which are arranged in two tiers. 
The nuclei of both tiers divide once more, and eventually the pro¬ 
embryo consists of four tiers of four cells each, the uppermost of which 
is open at the top. The lowest of the tiers becomes the embryo proper, 
the next tier above elongates to form the suspensor, and the uppermost 
of the completely walled tiers constitutes the rosette. 
The structure and development of the gametophytes and embryogeny 
of Pseudolarix follow the normal Abietinous type. 
LITERATURE CITED. 
Arnoldi, W. (’00) : Beitrage zur Morphologie der Gymnospermen, IV. Was sind die Keimblaschen 
oder ‘ Hofmeisters-Korperchen ’ in der Eizelle der Abietineen ? Flora, Bd. lxxxvii, 
1900 , p. 194 . 
Belajeff, W. (’93) : Zur Lehre von dem Pollenschlauche der Gymnospermen (zweite Mitteilung). 
Ber. d. Deutsch. Bot. Gesellsch., Bd. xi, 1893 , p. 196 . 
Blackman, V. H. (’98) : On the Cytological Features of Fertilization and Related Phenomena in 
Pinus sylvesiris , L. Phil. Trans. Roy. Soc., London, B, vol. cxc, 1898 , p. 395 . 
Cavara, F. (' 00 ) : Osservazioni morfologiche sulle Gimnosperme, I. Oogenesi nell’ Abies 
pectinata. Bull. Soc. Bot. Ital., 1900 , p. 317 . 
Chamberlain, C. J. (’ 99 ) : Oogenesis in Pinus Laricio. Bot. Gaz., vol. xxvii, 1899, p. 268. 
-(’ 06 ) : The Ovule and Female Gametophyte of Dioon. Bot. Gaz., vol. xlii, 
1906, p. 321. 
Coker, W. C. (’03): On the Gametophytes and Embryo of Taxodium. Bot. Gaz., vol. xxxvi, 
1903 , p. 1 . 
