1018 Tabor .— The Leaf Buds of Archytaea alternifolia. 
many Rosaceae, Saxifragaceae, &c., by Haberlandt ( 2 ), Reinke ( 7 ), von 
Minden ( 5 ), and others. 
There is, however, one important point of distinction, and that is that, 
instead of the water escaping by a well-defined pore between two epidermal 
cells of special construction, or by a group of such pores, it finds its exit 
by numerous small intercellular spaces. These are found between all the 
epidermal cells of the upper two-thirds of the tooth (PI, LXXIX, Figs. 5 and 8). 
At the base of the tooth no intercellular spaces are found between the 
epidermal cells or the two or three subjacent layers. These cells, moreover, 
have thicker cell-walls (PL LXXIX, Fig. 5), and sometimes show indication 
of divisions (PL LXXIX, Fig. 7). 
It is these cells, forming a shallow plinth for the tooth, which later 
acquire the dark-coloured contents already referred to. They are able 
to resist the fate which overtakes the thin-walled cells of the tooth, when 
freely exposed to desiccation by the later growth of the leaf. 
Solereder (8, page 133) describes Camellia as having leaf teeth which 
1 figure as secretory organs . . . they contain the termination of a vascular 
bundle in connexion with an epithemoid tissue, and above the latter 
a secretory epidermis of palisade-like radially elongated cells \ 
This probably refers to an observation of Reinke’s ( 7 ), who described 
the structure of the leaf teeth in Camellia japonica , and considered the 
epidermis of the tooth as a glandular tissue, which excreted a mucilaginous 
secretion through the cuticle. 
It was unfortunately impossible to confirm this observation; the only 
material of Camellia available was in a condition too advanced to show 
functional leaf teeth. In some shoots of The a viridis , kindly supplied by 
Messrs. Veitch, the young leaves showed prominent teeth with a clear 
hyaline, viscid apical portion. This terminal portion of the tooth consisted 
of a central core of cells, covered externally by an epidermis of palisade-like 
cells ; the whole structure strongly resembling the glandular ‘ villi ’ figured 
by Hanstein ( 3 ), Groom ( 1 ), and others, and obviously corresponding to the 
teeth described by Reinke ( 7 ) in Camellia . No excretion of water was 
observed when a shoot was placed under a pressure of 60 cm. of mercury 
for four hours, but no definite conclusion can be drawn from this isolated 
observation. 
Similar teeth have been described and figured by Virchow (12) and 
Tschirch ( 11 ) for Thea sinensis. In The a and Camellia the teeth—even if 
they should function as hydathodes, which does not seem probable—cannot 
subserve the same functions as those of Archytaea. The buds are small, 
and the shoot elongates rapidly, so that the young leaves are early 
separated, and the teeth are apparently functional on leaves which no 
longer invest the bud. 
The liquid which fills the space within the bud of Archytaea has been 
