BRACHIOPODA. 
137 
nal septa, it agrees throughout with Anoplotheca and C(elospira. In the 
original diagnosis of Leptoc(ELIA, L. flabellites was described and figured as pos¬ 
sessing a terebratuloid loop. It was, however, distinctly stated that the evi¬ 
dence of this structure was confined to a single specimen containing cavities in 
its filling of quartz, which corresponded to the restoration given. Subsequent 
investigations have not corroborated this observation. 
The specimens of this species are not favorably preserved for the retention 
of the brachial apparatus, those from Cumberland, Maryland, being replaced 
by silica and often filled with coarsely crystallized quartz, while those from 
New York, Gaspe and the South American localities are usually in the form 
of casts in an arenaceous sediment. Later observations have not shown any 
trace of the organ described, and it is highly improbable that a species agreeing 
in all known points of structure with the spirigerous groups just discussed, and 
having also a fibrous shell structure, should possess a terebratuloid loop. Ref¬ 
erence has been made to the fact that in Codospira concava the stout umbonal 
blades of the primary lamellae and the loop are frequently all that is retained 
of the brachial apparatus, the rest of the brachidium being very delicate ; 
the parts thus retained are by themselves certainly suggestive of terebratuloid 
structure, and not unlike the loop ascribed to L. flabellites. With our present 
knowledge it would be unwise to separate Lrptoccelia too widely from 
Anoplotheca and Ccblospira. 
There are but two other species which can safely be referred to the group 
of L. flabellites^ namely, L. dichoioma and L. flmbriata, Hall, also from the 
Oriskany fauna at Cumberland. The latter shell possesses a peculiarity in the 
extension, from between the cardinal extremities of the valves, of the inner 
lamina of the shell substance in the form of a row of spinules having the 
appearance of matted cilia. (See Plate LIII, figs. 47, 50.) 
Leptomlia flabellites, the type species, is remarkable for its wide distribution. 
Like Tropidoleptus carinatus and Vitulina pustulosa, it abounds in the lower 
Devonian strata of South America. Morris and Sharpe described the species 
under the name Atrypa palmata, from material collected by Darwin in the 
Falkland Islands; Salter described it as Orthis Aymara, from various localities 
