278 Wisconsin Academy of Sciences, Arts, and Letters. 
homogeneous in structure but consist of numerous smaller granu¬ 
les. The linin on the other hand shows scarcely if any granular 
structure. 
Although it is evident that the chromosomes of the equatorial 
plate are double structures, the halves always lie close together 
and are never separated in the way as has been described for the 
chromosomes of certain other plants. In no case have I observed 
a transverse segmentation of the chromosomes at this or any other 
stage. 
As mentioned above, the spindle fibers may be attached at the 
middle or at or near one end. In the case the attachment is near 
one end, each half as it is drawn toward the pole assumes the form 
of a J as is shown in figure 2 and in the chromosome at the right 
of figure 3. When the attachment is at one end, more or less 
straight rods pass to each pole. This condition is evident from 
the anaphasic chromosome in the center of figure 3. Sometimes 
the attachment of fibers is about the middle of the chromosomes, 
in which case Y-shaped figures result. This condition is repre¬ 
sented at the left in figure 3. As the anaphasic chromosomes are 
passing to the poles, they are quite separate from one another and 
do not mingle very closely until the poles are reached and then 
not so closely as described by some authors. It is noteworthy 
that the anaphasic chromosomes, being in the forms of Y’s, J’s, 
or I’s, are easily distinguishable from one another. I have ob¬ 
served that there is a tendency for the different forms habitually 
to pass in pairs toward the poles. In figure 5 three pairs of 
chromosomes may be seen at one pole, two Y-shaped chromosomes 
at the left, two more or less straight rod-shaped ones at the right 
and two J-shaped ones in the center. 
Chromosomes, occurring in pairs on the spindle, have been re¬ 
ported by Strasburger, (’05, ’075, ’08, ’ll), and more recently 
by Muller (’09, T2) and Gates (’12). That chromosomes occur 
in pairs in resting somatic nuclei is also apparent in many plants 
containing prochromosomes, as I have previously reported (Over- 
ton, ’05, ’09a). Lundegard (’10c) denies the existence of such 
a pairing in Allium and Yicia. Sharp (’13) fails to find such 
pairing in Yicia, and von Schustow (T3) denies its existence in 
Allium. 
That the metaphasic and anaphasic chromosomes are not homo¬ 
geneous, but consist of two stainable substances, is evident from 
figures 2 and 3. During the passage of the chromosomes from the 
