306 Wisconsin Academy of Sciences, Arts, and Letters. 
distinctly visible in many cases in the resting nucleus. The fact 
that in this plant a chromosome may have its own nueleole seems 
to support the persistence of karyomere-like structures in the 
resting reticulum although the membranes of the karyomeres 
may be broken down by the inosculation of neighboring vacuoles. 
Sharp (’13) points out that in certain resting nuclei of Yieia 
heavier bands are present which represent reticulate chromosomes, 
but he observed them joined by fine anastomoses to form a con¬ 
tinuous net. It appears to me that these fine anastomoses repre¬ 
sent only the boundaries of inosculating vacuoles of the chromo¬ 
some bands. 
Lundegard ('12 c) fails to find in Allium any definite orienta¬ 
tion of the chromosome bands; Sharp (’13) holds that the reticu¬ 
late bands of Yieia represent the chromosomes of the telophases, 
and figures such bands as more or less polarized and visible dur¬ 
ing rest; Fraser and Snell ('ll) and Lundegard ('09, '10a, '12c) 
state that the chromosomes of Vicia become indistinguishable 
during rest. Sakamura ('14) figures the chromosome bands of 
the telophases in Vicia cracca, as they become alveolated to form 
the resting nucleus, as remaining more or less polarized and states 
that these alveolated chromosomes are distinct during rest. Digby 
('10), Fraser and Snell ('ll), Beer (’12) and others have also 
described a polarity in the arrangement of the chromosomes. Mul¬ 
ler (’12) states that in rapidly growing merismatic regions of 
Najas, often the telophasic structures are carried over into the 
next succeeding prophase. Additional evidence of the persistence 
of the chromosomes during rest has been brought out by the ob¬ 
servations of Gregoire and his students, and the work of other ob¬ 
servers tends to show that the loss of chromosome identity dur¬ 
ing the reticulated condition is only apparent. The study of the 
formation of a finely divided reticulum in the telophases and its 
behavior in the prophases has led in many cases to the identifica¬ 
tion of the reticulated chromosomes during rest. 
Many authors have recently come to the conclusion that a con¬ 
tinuous chromatic spirem does not exist either at prophase or at 
telophase. Strasburger ('05) was unable to find such a spirem 
in the somatic cells of Funkia and Galtonia. Gregoire and Wy- 
gaerts (’03) do not figure a long, thin spirem in Trillium. Gre¬ 
goire (’06) figures a long slender spirem in Allium but holds that 
it is not continuously chromatic. Gregoire and his students are 
of the opinion that a continuously chromatic spirem does not ex- 
