22 
D.J. Kitchener, L.H. Schmitt, P. Strano, A. Wheeler, A. Suyanto 
Supraorbital length 
Figure 9 Plot of nasal inflation breadth, NIB, versus supraorbital length, NIL, for the taxa Rhinolophus simplex simplex, 
R. s. parvus, R. s. subsp. nov. R. megaphyllus (□) and R. borneensis parvus. The oblique broken line joins 
values where NIB = NIL. Other taxon symbols as for Figure 1, caption. Following Hill (1992), those forms 
with values generally below the line should be R. megaphyllus while those generally above the line are other 
species in the ferrumequinum' group. 
4a, b). Future collections from Tanimbar and Wetar 
Is may help clarify its taxonomic status with 
respect to R. s. simplex. 
Rhinolophus simplex parvus Goodwin, 1979 
Rhinolophus borneensis parvus Goodwin, 1979:102- 
105. 
Holotype 
American Museum, Natural History No. 237766, 
adult male, skin and skull, collected 27 March 1968. 
Type locality 
Lia Hoo Cave, nr Fatu Maca village, 11 km S 
Baucau, Timor, altitude ca. 550 m. 
Diagnosis 
Rhinolophus simplex parvus differs from 
Rhinolophus s. simplex and R. s. keyensis as 
diagnosed earlier for these subspecies. 
Differs from Rhinolophus simplex subsp. nov. in 
averaging larger (but with measurements 
overlapping) in all skull and dentary characters 
except supraorbital length and lower tooth row 
length (Table la). Also the relationship between 
lower tooth row length and : nasal inflation 
breadth, C'M 3 length and outer M 3 M 3 width differs 
(Figures 7a,b,c). General body measurements also 
average larger except those related to facial 
foliations: maximum noseleaf breadth, basal sella 
length, vertical sella height, maximum vertical sella 
breadth. 
Description 
Apart from differences described in the 
diagnoses of R. s. simplex, R. s. parvus is very 
similar to that subspecies. The skull, however, 
tends to have the juncture of supraorbital and 
lambdoidal ridges level with orbital cavity mid 
point (36%), just posterior (36%) or well posterior 
from that point (28%); anterior sphenoid/ 
pterygoid bridge tends to obscure more of 
sphenorbital sinus when viewed from ventral 
aspect. 
Pelage and skin colour also differs slightly. 
Dorsal pelage Cinnamon Brown, which is colour of 
distal one-third of hairs, basal part of which is 
Drab. Ventral surface pure Fawn Color. Wings 
Grayish Brown, Ears Fawn Color. 
