Morphological variation in Pteropus lombocensis 65 
Table 2 Multiple regressions on sex and island populations (Lombok, Flores, Lembata and Alor) of Pteropus 
lombocensis for skull, dentary, dental and external body characters. F values are presented for the main 
effects and their interactions. For explanation of character codes see Material and Methods section. 
Significance levels are *, 0.05>P>0.01; “, 0.01>P>0.001; ***, P<0.001. 
CHARACTER 
MAIN EFFECTS 
SEX ISLAND 
INTERACTION 
SEX. ISLAND 
GSL 
1.319 
1.455 
0.079 
CBL 
0.750 
2.082 
0.064 
PL 
0.982 
1.819 
0.064 
MFW 
0.881 
1.705 
0.509 
RL 
0.290 
0.466 
0.339 
IOB 
2.596 
0.870 
0.533 
ZW 
0.176 
0.697 
1.967 
BB 
0.497 
0.670 
1.834 
DL 
1.582 
3.028 
0.581 
C'C' 
1.346 
0.865 
1.368 
p4p4 
0.002 
1.589 
1.924 
M'M' 
0.047 
1.255 
0.150 
C'M 2 
0.044 
10.788*“ 
1.513 
c,m 3 
4.902* 
44.522*** 
2.232 
P 3 L 
1.605 
0.853 
1.772 
P 3 B 
2.178 
8.942“ 
0.104 
M'L 
0.108 
1.382 
0.926 
M'B 
0.997 
12.537“* 
0.677 
FA 
0.003 
1.846 
1.544 
D2M 
0.020 
1.436 
1.334 
D3M 
0.020 
1.266 
0.515 
D3P1 
0.504 
2.116 
1.288 
D3P2 
0.304 
0.862 
2.765 
D4M 
0.251 
1.334 
1.598 
D5M 
0.278 
1.144 
1.387 
TIB 
1.531 
4.417* 
6.046** 
PES 
3.685 
1.877 
2.023 
EAR 
0.098 
2.446 
0.626 
Degrees of freedom 
1,17 
3,17 
3,17 
Outside c'-C' width 
Figure 4 Plot of upper maxillary tooth row length 
versus width outside C'C 1 . 
SYSTEMATICS 
Pteropus lombocensis lombocensis Dobson, 1878 
Pteropus lombocensis Dobson, 1878: 34, pi. iii, fig. 2, 
pi. iv, fig. 4. 
Holotype 
Natural History Museum, London, Number 
64.4.12.2, adult male, 'cabinet skin' and skull. 
Type locality 
Lombok Island, Nusa Tenggara. 
Specimens examined 
Lombok Island (all adults) 
Desa Pelangan, 8°48'S, 115°56 E, Id, WAM 
M30603; Desa Kuta, 8°55'S, 116°15'E, 3dd 15, 
WAM M30611-4; Matahari Inn, Near Kuta, 4d d 
25 2, WAM M(36108, 36155, 36158, 36172, 36174- 
5). 
Sumbawa Island (all subadults) 
Desa Daha, 8°45’S, 118°26'E, 3dd 15, WAM 
M(31710-l, 31722, 31742). 
