Morphological variation in Pteropus lombocensis 
1908. Pteropus g. griseus occurs on Wetar Island (un¬ 
published record), immediately to the east of Alor 
Island, and some other islands in the gondwanic 
outer Banda Arc (Timor and Semau Island 
[Goodwin 1979] and Roti Island [unpublished 
record], A larger form of P. alecto, probably P. 
aledo gouldi Peters, 1867, also occurs on Timor Is¬ 
land [unpublished record]). Pteropus alecto morio 
occurs on Sumba and Savu Islands (Hill in Corbet 
and Hill 1992; unpublished records). Pteropus 
vampyrus occurs on two outer Banda Arc islands, 
Timor (Goodwin 1979) and Roti (unpublished re¬ 
port). 
In conclusion, the small Pteropus species in Nusa 
Tenggara (P. lombocensis, P. g. griseus, P. alecto 
morio ) show a pattern of distribution that is 
repeated by a number of other bat taxa. That is, a 
species that is widely distributed in the inner 
Banda Arc either differentiates morphologically 
into subspecies in the outer Banda Arc (Sumba, 
Savu, Roti, Semau, and Timor) or is replaced there 
by allied species (see Kitchener et al. 1994 a,b) 
ACKNOWLEDGEMENTS 
We gratefully acknowledge the support of both 
the current Director, Mr A. Reeves, and the 
previous Director, Mr J. Bannister, of the Western 
Australian Museum; Drs M. Amir, Director, 
Balitbang Zoologi (LIPI); and the Directors of the 
Department responsible for the conservation of 
wildlife in Nusa Tenggara Barat, Ir P. Supriadi and 
Nusa Tenggara Timor, Ir J. Mochtar. To our 
colleagues Dr R. How and Mr R. Johnstone, 
Western Australian Museum; Mr A. Gunnell, 
Curtin University of Technology; and Ir I. 
Maryanto and Drs A. Suyanto, Museum 
Zoologicum Bogoriense, who participated in the 
exhausting field work, we extend our thanks for 
both their effort and companionship. Dr How 
kindly reviewed the manuscript. 
Expedition costs were defrayed by grants to Dr 
D. Kitchener from: National Geographic Society, 
Washington, Australian National Parks and 
Wildlife Service and the Scott Trust, Sydney. 
Garuda Indonesia kindly defrayed freight costs of 
the expeditions. Skulls were prepared by Mr M. 
Salotti, Western Australian Museum and Mr 
Wayne Walters, Curtin University of Technology, 
67 
the latter also recorded all external measurements. 
Mrs N. Cooper assisted with the statistical analy¬ 
ses. Mrs A. Nevin, Western Australian Museum 
kindly typed the manuscript. 
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Manuscript received 10 January 1994; accepted 14 July 1994. 
