Systematic review of Nyctimene cephalotes and N. albiventer 
129 
Table 1 (continued) 
C'M 1 
IOB 
POB 
MFB 
p4p4 
ML 
CH 
c , m 2 
FA 
MC3 
MC4 
MC5 
TV 
EAR 
9 
9 
9 
9 
9 
9 
9 
9 
5 
5 
5 
5 
5 
5 
10.62 
6.44 
6.18 
5.18 
6.02 
23.79 
13.92 
11.96 
68.6 
49.1 
44.8 
48.6 
25.1 
16.2 
0.28 
0.40 
0.60 
0.13 
0.23 
0.41 
0.31 
0.25 
0.6 
1.9 
2.2 
2.7 
1.0 
0.3 
10.34 
6.03 
5.51 
4.96 
5.47 
23.20 
13.41 
11.66 
68.1 
47.4 
42.3 
45.2 
23.7 
15.8 
11.11 
7.17 
7.39 
5.37 
6.26 
24.37 
14.47 
12.41 
69.5 
51.8 
47.7 
52.0 
26.2 
16.5 
26 
26 
26 
25 
26 
26 
26 
26 
24 
24 
24 
24 
24 
24 
10.16 
5.65 
5.77 
5.16 
5.58 
22.55 
12.99 
11.46 
63.4 
45.1 
41.8 
44.6 
22.4 
14.2 
0.43 
0.35 
0.44 
0.20 
0.23 
0.67 
0.72 
0.44 
2.1 
1.6 
1.5 
1.7 
1.3 
0.6 
9.02 
4.94 
5.17 
4.79 
4.89 
21.02 
11.47 
10.21 
59.1 
41.4 
38.6 
40.2 
20.6 
12.9 
10.82 
6.13 
6.94 
5.61 
5.94 
23.69 
14.23 
12.09 
67.6 
48.0 
44.1 
47.2 
24.8 
15.2 
35 
35 
35 
34 
35 
35 
35 
35 
29 
29 
29 
29 
29 
29 
10.28 
5.85 
5.88 
5.16 
5.69 
22.87 
13.23 
11.59 
64.3 
45.8 
42.3 
45.3 
22.9 
14.5 
0.44 
0.50 
0.51 
0.19 
0.30 
0.82 
0.76 
0.46 
2.7 
2.2 
2.0 
2.4 
1.6 
1.0 
9.02 
4.94 
5.17 
4.79 
4.89 
21.02 
11.47 
10.21 
59.1 
41.4 
38.6 
40.2 
20.6 
12.9 
11.11 
7.17 
7.39 
5.61 
6.26 
24.37 
14.47 
12.41 
69.5 
51.8 
47.7 
52.0 
26.2 
16.5 
12 
12 
12 
12 
12 
12 
12 
12 
12 
12 
12 
12 
12 
12 
9.57 
5.69 
6.02 
4.76 
5.19 
21.49 
12.86 
10.84 
58.2 
42.2 
38.5 
40.6 
20.7 
15.0 
0.24 
0.24 
0.35 
0.17 
0.10 
0.51 
0.31 
0.26 
1.3 
1.1 
0.7 
1.0 
1.5 
0.5 
8.98. 
5.29 
5.21 
4.50 
5.03 
20.58 
12.33 
10.27 
55.1 
40.7 
37.5 
38.7 
18.6 
14.2 
9.80 
6.12 
6.44 
5.05 
5.38 
22.11 
13.32 
11.18 
59.7 
44.5 
39.6 
42.7 
24.1 
15.6 
44 
44 
44 
44 
44 
44 
44 
44 
44 
44 
44 
44 
44 
44 
10.01 
5.81 
5.66 
5.00 
5.60 
22.00 
13.38 
11.28 
60.1 
43.5 
39.6 
42.1 
21.3 
15.4 
0.33 
0.28 
0.41 
0.17 
0.19 
0.49 
0.47 
0.29 
1.2 
1.0 
1.1 
1.0 
1.6 
0.6 
8.39 
5.30 
5.03 
4.56 
5.24 
21.04 
12.34 
10.76 
58.0 
41.6 
37.5 
40.0 
18.0 
14.3 
10.50 
6.49 
6.65 
5.39 
5.97 
23.10 
14.46 
12.10 
63.0 
45.7 
43.1 
44.2 
25.4 
16.9 
56 
56 
56 
56 
56 
56 
54 
56 
56 
56 
56 
56 
56 
56 
9.91 
5.78 
5.74 
4.95 
5.51 
21.89 
13.26 
11.18 
59.7 
43.2 
39.4 
41.8 
21.2 
15.3 
0.36 
0.27 
0.42 
0.20 
0.24 
0.53 
0.49 
0.34 
1.4 
1.2 
1.1 
1.2 
1.6 
0.6 
8.39 
5.29 
5.03 
4.50 
5.03 
20.58 
12.33 
10.27 
55.1 
40.7 
37.5 
38.7 
18.0 
14.2 
10.50 
6.49 
6.65 
5.39 
5.97 
23.10 
14.46 
12.10 
63.0 
45.7 
43.1 
44.2 
25.4 
16.9 
N. cephalotes was fixed as Ambon by Andersen 
(1912). The above documentation of the 
morphological separation of the Sulawesi 
population from N. c. cephalotes warrants its 
description as a new subspecies of N. cephalotes. 
Nyctimene keasti populations 
DFA was run for all 23 characters for the islands 
Kai Besar, Kai Kecil, Selaru and Yamdena, with 
Banda Neira unallocated. A reduced set of five 
characters was selected from this above analysis. 
These were palatal width between the lingual 
aspect of P*P; ear length; forearm length; braincase 
breadth; minimum length from posterior margin of 
incisive foramen to palate posterior margin. This 
latter DFA produced two significant functions, 
which combined explained 98.4% of the variation 
(Figure 6). A total of 81.5% of individuals were 
correctly classified to their appropriate island, with 
misclassification occurring between the two 
Tanimbar islands. One of the unallocated Banda 
Neira specimens classified with the Selaru 
population and the other with the Yamdena 
population. 
The DFA run with 23 characters and only two a 
priori groups: Tanimbar group (Selaru and 
Yamdena) and Kai group (Kai Kecil and Kai Besar) 
produced a similar discriminant histogram to that 
produced from a subset of the five characters that 
were selected from the above analysis (palate 
breadth at P 4 P; incisive foramen to posterior palate 
length, PIF; braincase breadth, BB; postorbital 
breadth, POB; and forearm length, FA). The 
function extracted from this latter analysis was 
highly significant and resulted in all individuals 
being correctly allocated to their appropriate group 
(Figure 7). The characters loading heavily (>0.6) on 
this function were P*P, FA and BB (Table 5). 
The type locality of the nominate subspecies of 
N. keasti is Pulau Dullah, Kai Kecil. The above 
analysis documents clearly the morphological 
separation of the Tanimbar form from N. k. keasti. 
It indicates that the Tanimbar form warrants 
description as a new taxon. It also shows that some 
