132 
D.J. Kitchener, W.C. Packer, A. Suyanto 
6 -i 
4 - 
-4-2 0 2 4 
Function 1 
Figure 5 Canonical variate analysis between the 
Sulawesi population (Nyctimene cephalotes 
aplini subsp. nov.) (■) and those of the group 
of Seram, Ambon and Buru (N. c. cephalotes) 
(□). The histogram for function 1 was based 
on a selection of five skull characters, with 
males and females combined. 
subspecies below. 
The Aru form, is still phenetically close to the 
Papua New Guinea form, but is more 
differentiated morphologically from the Papua 
New Guinea form than was thought to be the case 
in Kitchener et al. (1993). Its subspecific status is 
not determined. Within the island groups of 
Tanimbar and Kai there has been considerable 
morphological differentiation between populations 
of subspecies of N. keasti inhabiting different 
islands. This has occurred to a lesser extent also 
between the Ambon and Seram populations of N. 
c. cephalotes. 
SYSTEMATICS 
Nyctimene cephalotes (Pallas, 1767) 
Diagnosis 
Nyctimene cephalotes differs from Nyctimene 
Table 5 Canonical variate function coefficients from 
DFA between populations of Nyctimene keasti 
keasti from the Kai Islands and N. k. tozeri 
from the Tanimbar Islands. Standardised 
values, followed by (in brackets) 
unstandardised values, for skull, dental and 
external body characters. For explanation of 
character codes see Materials and Methods 
section. 
CHARACTER 
FUNCTION 1 
P'P 1 
0.6922 ( 3.9545) 
FA 
-0.7606 (-1.9787) 
BB 
0.7532 (2.5839) 
POB 
-0.4283 (-1.0462) 
PIF 
0.3125 ( 0.2625) 
Constant 
-40.5598 
Table 6 Canonical variate function coefficients from 
DFA between populations of Nyctimene 
albiventer, N. a albmenter, N. a. papuanus and 
N. a. subsp. indet. from Halmahera group, 
and Papua New Guinea and Aru, 
respectively. Standardised values, followed by 
(in brackets) unstandardised values, for skull 
and dental characters. For explanation of 
character codes see Materials and Methods 
section. 
CHARACTER 
FUNCTION 1 
FUNCTION 2 
p4p4 
0.7335 (3.0491) 
0.4262 
(1.7717) 
C'M' 
0.1907 ( 0.7626) 
-0.9287 (-3.7137) 
RH 
-0.2628 (-0.8189) 
0.8977 
(-2.7979) 
MFW 
-0.5806 (-2.3344) 
0.1779 
(0.7151) 
GSL 
0.6701 (1.0692) 
0.1438 
(0.2295) 
Constant 
Variation 
-35.8486 
-3.3963 
explained (%) 
74.5 
25.5 
Figure 6 Canonical variate analysis among four island 
populations of Nyctimene keasti: Kai Besar, 
Kai Kecil (N. k. keasti), Selaru and Yamdena 
(N. k. tozeri subsp. nov.). The Banda Neira 
population was ungrouped. The DFA plots 
of functions 1 and 2 were based on a selection 
of five skull, dentary and external body 
characters, with males and females 
combined. Symbols as for Figure 2. 
