Function 1 
Systematic review of Nyctimene cephalotes and N. albiventer 
133 
Function I 
Figure 7 Canonical variate analysis between Nyctimene 
keasli populations from the Kai Island group 
(Kai Besar and Kai Kecil) (N. k. keasti) (■) 
and the Tanimbar Island group (Selaru and 
Yamdena) (N. k. tozeri subsp. nov.) (□). The 
Banda Neira population was ungrouped (0). 
The histogram for Function l was based on a 
selection of five characters (skull, dentary 
and external body), with males and females 
combined. 
Function 2 
Figure 8 Canonical variate analysis between Nyctimene 
albiventer populations from the Haimahera 
island group (Haimahera, Morotai and 
Ternate) (N. a. albiventer), Aru (N. a. subsp. 
indet.) and Papua New Guinea (N. a. 
papuanus). The DFA plots of functions 1 and 
2 were based on a selection of 5 characters 
(skull and dental), with males and females 
combined. Symbols as for Figure 2. 
Table 7 Canonical variate function coefficients from 
DFA between populations of Nyctimene 
albiventer from Aru islands and Papua New 
Guinea. Standardised values, followed by (in 
brackets) unstandardised values for skull, 
dental and external body characters. For 
explanation of character codes see Materials 
and Methods section. 
CHARACTER 
FUNCTION 1 
C'M' 
0.7909 ( 3.0302) 
RH 
-0.6888 (-2.1155) 
EAR 
0.5633 (0.6949) 
RL 
0.4382 ( 1.6215) 
IOB 
-0.4345 (-1.4417) 
Constant 
-23.1370 
albiventer by averaging larger in all measurements 
(Table 1). Incisive foramen to posterior palatal 
length shorter relative to postorbital breadth 
(Figure 10); C,M 2 length generally longer relative 
to M'M 1 length (Figure 11); and forearm longer 
relative to ear length (Figure 12). Female dorsum 
paler than that of males. 
It differs from Nyctimene keasti by averaging 
larger in all measurements except for cranial height 
(Table 1). Condylobasal length generally longer 
relative to both rostrum height, interorbital breadth 
and cranial height (Figure 13A,B,C, respectively); 
and forearm longer relative to ear length (Figure 
12 ). 
Nyctimene cephalotes aplini Kitchener, 
subsp. nov. 
Holotype 
Museum Zoologicum Bogoriense Number MZB 
15903; adult male; weight 46 g; carcase fixed in 
10% formalin and preserved in 75% ethanol; skull 
and mandibles separate; collected by Drs 
Augustine Suyanto on 28 July 1990. 
-4-2 0 2 
Function 1 
Figure 9 Canonical variate analysis between two 
populations of Nyctimene albiventer: Aru (N. 
a. aruensis subsp. nov.) (■) and Papua New 
Guinea (N. a. papuanus ) (□). The histogram 
for function 1 was based on a selection of 
five characters (skull, dental and external 
body), with males and females combined. 
