Incisiue Foramen-Post Palate Length 
134 
D.J. Kitchener, W.C. Packer, A. Suyanto 
Postorbital Breadth 
Figure 10 Plot of incisive foramen to posterior palate 
length versus postorbital breadth for all 
island populations of Nyctimene studied. 
Males and females combined. Symbols as for 
Figure 2. 
Paratypes 
Tornado, Lake Lindu area Central Sulawesi, 
WAM M27659 A 2, skin and skull; Soroako, 
Sulawesi Selatan, WAM M27676 A 2, skin and 
skull; WAM M33145, near Bontobonto, Sulawesi 
Selatan (c. 5°07'S, 119°39'E), A 2, spirit carcase and 
skull; Maros, Sulawesi Selatan (c. 4°59'S, 119°35'E), 
WAM (M33146, M33207), IS S skin and skull, 
WAM (M33208, M33210-12), 4 SS, spirit carcase 
and skull. 
Type locality 
Maros, Sulawesi Selatan (= S. Sulawesi) (c. 4°59', 
119°35'E), altitude c. 50 m; mist-netted. 
Diagnosis 
Nyctimene cephalotes aplini differs from N. c. 
cephalotes in averaging larger in all skull, mandible 
and external measurements; in some it is absolutely 
larger (Table 1). Condylobasal length greater 
relative to both mesopterygoid fossa breadth, MFB, 
and C'M 1 length (Figures 14A,B, respectively); and 
zygomatic width greater relative to MFB (Figure 
15). 
Description 
Skull, dentary and dentition 
Large skull with greatest length 31.7 ± 0.47, broad 
zygomatic width 21.4 ± 0.48, palate long and broad 
both anteriorly, with C*C breadth 5.8 ± 0.27, and 
posteriorly, with M'M 1 breadth 9.6 ± 0.19; rostrum 
short, nasal dorsal surface almost horizontal, 
occasionally with dorsal distal inclination; rostrum 
high; premaxillary part of nares terminates directly 
above incisors or projects slightly anterior to them; 
braincase typical of N. cephalotes with cranium 
inflated to maximum height at junction of strong 
sagittal and weak postorbital crests; postorbital 
breadth large; usually deeply basined and opens 
anteriorly into deep frontal sulcus; frontals 
converging slightly posteriorly; sagittal and 
lambdoidal crests moderately high; incisive 
foramen subcircular, reach posteriorly almost to 
line joining P 2 P 2 anterior face; posterior palate 
extends well beyond M 1 , its margin a broad U 
shape; basi- and presphenoid median ridge 
prominent; pterygoid process low and slightly 
curved ventrally and laterally. 
Upper tooth row slightly curved; upper incisors 
in contact, sited close to line joining C'C 1 
Figure 11 Plot of CjMj length versus outside M'M 1 
breadth for all island population of 
Nyctimene studied. Males and females 
combined. Symbols as for Figure 2. 
