138 
D.J. Kitchener, W.C. Packer, A. Suyanto 
Mesopterygoid Fossa Breadth 
Figure 15 Plot of zygomatic width versus 
mesopterygoid fossa breadth for individual 
Nyctimene cephalotes. Symbols as for Figure 
2 . 
ventral surface a uniform Cream Color. Patagia in 
males Olive Brown; in females a paler Fawn Color. 
Spots on patagia spectrum Yellow and distributed 
similarly to N. cephalotes aplini. 
Distribution 
Seram, Ambon and Buru Islands, Maluku 
Tenggara, Indonesia. It is unlikely that the 
Nyctimene from Timor Island attributed to N. 
cephalotes by Anderson (1912), Goodwin (1979) and 
Hill in Corbet and Hill (1992) is of that species. It is 
more likely to be associated with N. keasti. 
Nyctimene keasti Kitchener, 1993 
Nyctimene alhiventer keasti Kitchener et al., 1993: 
408-412 
Diagnosis 
Nyctimene keasti differs from N. cephalotes as 
described in the earlier diagnosis of N. cephalotes. It 
differs from N. alhiventer in averaging larger in all 
skull, mandible and external measurements. For 
example, greatest skull length 29.34 (28.04-30.78) 
56 v. 26.98 (24.67-28.45) 45; braincase breadth 12.64 
(11.64-13.94) 56 V. 11.93 (11.10-13.06) 45; zygomatic 
width 19.28 (18.11-20.40) 56 v. 18.07 (16.55-19.69) 
45; C'M 1 length 9.91 (8.39-10.50) 56 v. 8.82 (8.20- 
9.51) 45; forearm length 59.7 (55.1-63.0) 56 v. 54.9 
(49.9-59.2) 48; and ear length 15.3 (14.2-16.9) 56 v. 
13.2 (11.7-14.7) 48; forearm length generally longer 
relative to ear length (Figure 12); incisive foramen 
to posterior palate length, PIF, relatively greater 
relative to postorbital breadth (Figure 10); C,M 2 
length greater relative to M'M' width (Figure 11). 
Nyctimene keasti tozeri Kitchener, subsp. nov. 
Holotype 
Museum Zoologicum Bogoriense Number MZB 
15904, adult male, weight 29.5 gm, carcase fixed in 
10% formalin and preserved in 75% ethanol; skull 
and mandibles separate, liver preserved in 
ultrafreeze at WAM, collected by D.J. Kitchener 
and R.A. How on 27 April 1993. 
Type locality 
Lorulun (Figure 1) c. 20 km N Saumlaki, 
Yamdena Island, Tanimbar group; Maluku 
Tenggara, Indonesia (7°52'S, 131°25'E); altitude 200 
m. Mist netted at c. 2m height in disturbed vine 
forest with the following genera of trees prominent: 
Ficus, Erythrina, Podocarpus, Alhizia as well as 
'Kenari' trees. The understory was mostly Lantana. 
There were numerous vines and creepers (see 
Kitchener and Maryanto 1994: Figure 5). 
Paratypes 
Yamdena Island: Desa Sifnana Omele, 7°56’S, 
131°18'E, WAM44398-401 (3 9 2, Id); Desa Lorulun 
(7°52'S, 131°25’E), WAM M(43776, 43782-88, 4394- 
95,43798,43801-06,43809) (102 2, 8d d); Saumlaki, 
7°59'S, 131°22'E, WAM M(43511-12, 43555-58, 
43564-66, 43600) (82 9, Id). Selaru Island: Desa 
Adaut, 8°09'S, 131°08’E, WAM M(44201-O5, M4408, 
M44300-03, M44310-11, M44316) (112 2,2d d). 
Diagnosis 
Nyctimene keasti tozeri differs from N. k. keasti in 
averaging larger in all skull, mandible and external 
measurements, except for incisive foramen to 
posterior palate length (PIF) and postorbital 
breadth (Table 1). Palatal length between P 4 P J 
lingual surface broader relative to both postorbital 
breadth and PIF (Figures 16 A,B, respectively); 
braincase breadth generally larger relative to 
postorbital breadth (Figure 17). It differs also from 
N. k. keasti in females having pelage of dorsum a 
similar dark Drab colour to that of males rather 
than females with a dorsum of Buff Yellow which 
is paler than the male Fawn color dorsum. 
