Systematic review of Nyctimene cephalotes and N. albiventer 
139 
B Incisiue Foramen-Post Palate Length 
Figure 16 Plot of palate breadth at P 4 P 4 versus (A) postorbital breadth and (B) incisive foramen to posterior palate 
length for individual Nyctimene keasti. Symbols as for Figure 2. 
Description 
Skull, dentary and dentition 
Intermediate sized skull with greatest length 
29.5±0.56; zygomatic width 19.4±0.45; braincase 
breadth 12.7±0.29 and C'M 1 length 10.0±0.34 (Table 
1). Very similar in shape to Nyctimene cephalotes 
aplini except that the junction of the postorbital 
ridges and sagittal crest closer to postorbital 
process base and frontal area immediately anterior 
to this junction is less basined than is the case in N. 
c. aplini. 
Externals 
Forearm moderately long 60.1+1.12; ear 
moderately long 15.4+0.60. wing membrane 
usually attached to pes between phalanx 1 of digits 
2 and 3 (56%, N=25); also commonly attached to 
phalanx 1 of digit 2 (16%, N=7), and phalanx 1 of 
digit 3 (27%, N=12) and rarely to between digits 3 
and 4 at level of phalanx 1 (2%, N=l). 
Pelage and patagia colouration very similar to N. 
c. cephalotes. Namely male and female dorsal 
surface of similar colour but with females having a 
paler head, face and ventral surface. 
Distribution 
Yamdena, Salaru and Larat islands, Tanimbar 
group, Maluku Tenggara, Indonesia. 
Etymology 
Named after Mr Tom Tozer, a distinguished 
retired gentleman, who for many years provided 
honorary assistance in the mammal section. 
Western Australian Museum, primarily with the 
curation of the Indonesian collection. 
Referred specimens 
The two specimens from Banda Neira Island, 
Band Group were included in the paratype series 
of Nyctimene albiventer keasti Kitchener, 1993 in 
Kitchener et al. (1993). These two specimens are 
treated as unallocated in the preceeding 
multivariate analysis because they are not readily 
attributable to either N. cephalotes or N. keasti. They 
may represent a distinct form of Nyctimene but if 
referrable to a recognisable form of keasti, they are 
clearly associated with N. k. tozeri. The taxonomic 
relationships of the Banda Island form will be 
clarified when more specimens become available 
for study. 
The specimen WAM M34549, adult male, alcohol 
and skull specimen, collected from Solea, Seram, 
was included in all the preceeding statistical 
analysis as an unallocated specimen. It has a short 
forearm (58.0) and skull and external 
measurements similar to N. keasti tozeri. It was 
clearly allocated to this taxon in the DFA. 
