140 
D.J. Kitchener, W.C. Packer, A. Suyanto 
Figure 17 Plot of braincase breadth versus postorbital 
breadth for individual Nyctimene keasti. 
Symbols as for Figure 2. 
Nyctimene albiventer albiventer (Gray, 1863) 
Cynopterus albiventer Gray, 1863: 261-63 
Holotype 
The Natural History' Museum, London Number 
BMNH 62.10.21.5; adult, 'cabinet' skin and skull, 
collected by Dr. A.R. Wallace in 1861. 
Type locality 
Morotai Island, Maluku Utara, Indonesia (c. 
2°30'N, 128°30'E) 
Specimens examined 
Listed in Kitchener et al. (1993) 
Diagnosis 
Nyctimene albiventer albiventer differs from N. a. 
papuanus in averaging smaller in all measurements 
except rostrum length (see Kitchener et al. 1993: 
Table 1). Forearm length generally smaller relative 
to ear length (Kitchener et al. 1993: Fig 4) and 
palatal breadth at FP 4 generally smaller relative to 
rostrum length (Kitchener et al. 1993: Figure 3b). 
If differs from N. albiventer subsp. indet. from 
Aru Island in averaging smaller in all 
measurements, except rostrum length, 
mesopterygoid fossa breadth and snout to vent 
length (see Kitchener et al. 1993: Table 1). P 4 P 4 
breadth also smaller relative to greatest skull length 
(see Kitchener et al. 1993: Figure 3a); and ear length 
longer relative to forearm length (see Kitchener et 
al. 1993: Figure 4). 
Description 
A small subspecies of N. albiventer with greatest 
skull length 25.68 ± 0.60 and forearm length 51.2 ± 
1.2 (see Kitchener et al. 1993, Table 1). The overall 
shape of skull, dentary, and dentition similar to N. 
cephalotes aplini. Its pelage and colour are similar to 
that described earlier for N. c. cephalotes. Wing 
membrane attached to pes on phalanx 1 of digit 3 
(Andersen 1912: 701). 
Distribution 
Morotai, Gilolo, Ternate and Goal area, N. 
Halmahera Island. 
Nyctimene albiventer papuanus K. Andersen, 1910 
Nyctimene papuanus K. Andersen, 1910: 621. 
Holotype 
The Natural History Museum, London Number 
Figure 18 Plot of rostrum height versus C'M 1 length for 
individual Nyctimene albiventer subsp. indet. 
and N. a papuanus. Symbols as for Figure 2. 
