192 
D.J. Kitchener, N. Cooper, I. Maryanto 
Figure 1 Locality of Myotis specimens examined in this study. 
this form is very decidedly smaller than M. 
adversus macropus (from South Australia). Tate 
(1952) examined three specimens of Myotis 
(Leuconoe) from Cairns, Northern Queensland, 
Australia, and although they were of similar size to 
Myotis adversus moluccarum from New Guinea, he 
considered them representative of M. adversus 
macropus, rather than M. a. moluccarum. 
Koopman (1984:12) also considered that only one 
subspecies of Myotis adversus (macropus) occurred 
in Australia and stated that its distribution 
extended from "northeastern Western Australia 
around the northern and eastern coasts to 
southeastern South Australia but apparently 
nowhere extending far inland" Richards (1983) and 
Mahoney and Walton (1988) recognised M. a. 
moluccarum from Northern Australia and M. a. 
macropus from south and eastern Australia. 
A number of terrestrial vertebrate faunal surveys 
in Nusa Tenggara and the Maluku Tenggara 
regions of Indonesia, between November 1987 and 
November 1993, were carried out jointly by staff of 
the Western Australian Museum and Museum 
Zoologicum Bogoriense. These surveys resulted in 
extensive collections of Myotis adversus (sensu lato) 
on islands from which they had not previously 
been recorded. These recent collections bridge the 
previous distributional gap between the Oriental 
and Australian M. adversus (sensu lato) and allow 
for a reappraisal of the taxonomy of some of the 
forms of M. adversus. 
This paper reports on a taxonomic reappraisal of 
Myotis adversus (sensu lato), based on a 
morphological examination of specimens 
principally from eastern Indonesia and Australia. 
MATERIALS AND METHODS 
A total of 149 specimens (listed in the specimens 
examined section) was examined from a number of 
localities in the Indonesian and Australo-papuan 
region (Figure 1). These were from Java (2), Nusa 
Penida (6), Kangean Island (1), Sumbawa Island 
(5), Moyo Island (1), Flores Island (3), Lembata 
Island (1), Pantar Island (1), Alor Island (38), Wetar 
Island (18), Yamdena Island (8), Seram Island (1), 
Timor Island (1), Savu Island (4), Papua New 
Guinea (2), Solomon Islands (2), Queensland (12), 
Northern Territory (2), Western Australia (22), 
New South Wales (1), Victoria (14) and South 
Australia (4). All the specimens from Indonesia are 
currently lodged in the Western Australian 
