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D.J. Kitchener, N. Cooper, I. Maryanto 
Table 2 Measurements, in mm, for (a) skull, dentary and dental characters, and (b) external characters (see Materials 
and Methods for explanation of character codes) of adult Myotis adversus (all populations); M. a. adversus 
(Java I; Kangean Is, Nusa Penida I., Sumbawa I., Moyo I., Flores I., Lembata 1., Pantar I., Alor I., Timor I., 
Savu I.); M. a. tanimbarensis subsp. nov. (Yamdena I.); M. a. ivetarensis subsp. nov. (Wetar I.); Myotis adversus 
subsp. indet. (New South Wales); Myotis moluccarum (all populations); M. m. moluccarum (Western Australia, 
Papua New Guinea, Seram I.); M. m. richardsi subsp. nov. (Queensland, Northern Territory)); M. moluccarum 
(Solomon Is) and Myotis macropus (Victoria, South Australia). N = sample size; SD = standard deviation; 
MIN = minimum; and MAX = maximum. 
Table 2a 
Taxon 
GSL 
CBL 
BB 
ZW 
MW 
CH 
RL 
AOB 
LIB 
Myotis adversus 
N 
73 
73 
73 
63 
73 
73 
73 
73 
73 
(all populations) 
MEAN 
16.55 
15.13 
8.11 
10.40 
8.56 
6.35 
4.85 
4.48 
4.06 
SD 
0.30 
0.30 
0.13 
0.21 
0.17 
0.13 
0.17 
0.14 
0.10 
MIN 
15.72 
14.47 
7.70 
9.60 
8.08 
5.89 
4.34 
4.12 
3.85 
MAX 
17.20 
15.85 
8.40 
10.75 
8.91 
6.64 
5.22 
4.76 
4.31 
Myotis adversus 
N 
46 
46 
46 
45 
46 
46 
46 
46 
46 
adversus 
MEAN 
16.58 
15.10 
8.14 
10.43 
8.58 
6.36 
4.85 
4.44 
4.06 
SD 
0.25 
0.25 
0.12 
0.18 
0.17 
0.13 
0.14 
0.11 
0.11 
MIN 
15.79 
14.59 
7.87 
9.81 
8.19 
6.06 
4.50 
4.12 
3.85 
MAX 
17.13 
15.68 
8.40 
10.75 
8.91 
6.64 
5.13 
4.72 
4.31 
Myotis adversus 
N 
8 
8 
8 
3 
8 
8 
8 
8 
8 
tanimbarensis 
MEAN 
16.14 
14.79 
8.07 
10.07 
8.29 
6.22 
4.65 
4.45 
4.06 
SD 
0.28 
0.27 
0.17 
0.40 
0.13 
0.14 
0.21 
0.12 
0.11 
MIN 
15.72 
14.47 
7.70 
9.60 
8.08 
5.89 
4.34 
4.32 
3.87 
MAX 
16.66 
15.31 
8.27 
10.30 
8.52 
6.35 
4.92 
4.66 
4.08 
Myotis adversus 
N 
18 
18 
18 
14 
18 
18 
18 
18 
18 
wetarensis 
MEAN 
16.71 
15.37 
8.04 
10.43 
8.62 
6.38 
4.94 
4.61 
4.12 
SD 
0.23 
0.24 
0.11 
0.11 
0.09 
0.11 
0.15 
0.16 
0.07 
MIN 
16.34 
14.92 
7.81 
10.21 
8.49 
6.20 
4.66 
4.16 
4.01 
MAX 
17.20 
15.85 
8.25 
10.62 
8.83 
6.61 
5.22 
4.76 
4.26 
Myotis adversus 
N=1 
15.94 
14.64 
8.16 
9.74 
8.53 
6.25 
4.67 
4.48 
3.97 
subsp. indet 
Myotis moluccarum 
N 
37 
37 
37 
34 
37 
37 
37 
37 
37 
(all populations) 
MEAN 
15.64 
14.48 
7.81 
10.06 
8.24 
6.08 
4.46 
4.22 
3.85 
SD 
0.34 
0.38 
0.18 
0.30 
0.24 
0.18 
0.17 
0.14 
0.12 
MIN 
14.98 
13.80 
7.48 
9.50 
7.90 
5.82 
4.17 
3.89 
3.53 
MAX 
16.37 
15.39 
8.19 
10.76 
8.91 
6.59 
4.92 
4.51 
4.20 
Myotis moluccarum 
N 
22 
22 
22 
22 
22 
22 
22 
22 
22 
moluccarum 
MEAN 
15.48 
14.29 
7.70 
9.92 
8.13 
6.01 
4.36 
4.16 
3.87 
SD 
0.23 
0.19 
0.10 
0.20 
0.15 
0.13 
0.11 
0.08 
0.09 
MIN 
14.98 
13.97 
7.48 
9.50 
7.90 
5.84 
4.17 
4.00 
3.71 
MAX 
15.82 
14.59 
7.86 
10.28 
8.41 
6.32 
4.62 
4.32 
4.07 
Myotis moluccarum 
N 
13 
13 
13 
13 
13 
13 
13 
13 
13 
richardsi 
MEAN 
15.95 
14.87 
8.01 
10.32 
8.49 
6.17 
4.62 
4.36 
3.88 
SD 
0.31 
0.35 
0.11 
0.28 
0.23 
0.23 
0.15 
0.09 
0.13 
MIN 
15.39 
14.27 
7.85 
9.74 
8.12 
5.82 
4.38 
4.21 
3.71 
MAX 
16.37 
15.39 
8.19 
10.76 
8.91 
6.59 
4.92 
4.51 
4.20 
Myotis moluccarum 
N 
2 
2 
2 
2 
2 
2 
2 
2 
2 
(Solomon Is) 
MEAN 
15.33 
14.00 
7.69 
9.75 
8.01 
6.10 
4.37 
3.91 
3.64 
SD 
0.36 
0.28 
0.18 
0.17 
0.11 
0.04 
0.09 
0.03 
0.06 
MIN 
15.07 
13.80 
7.56 
9.63 
7.93 
6.07 
4.30 
3.89 
3.59 
MAX 
15.58 
14.20 
7.82 
9.87 
8.09 
6.12 
4.43 
3.93 
3.68 
Myotis macropus 
N 
15 
15 
15 
12 
15 
15 
15 
15 
15 
MEAN 
16.75 
15.66 
8.36 
10.70 
8.70 
6.46 
4.84 
4.58 
4.05 
SD 
0.46 
0.46 
0.17 
0.38 
0.18 
0.22 
0.21 
0.21 
0.11 
MIN 
16.18 
14.91 
8.14 
10.20 
8.48 
6.19 
4.51 
4.32 
3.88 
MAX 
17.76 
16.58 
8.70 
11.42 
9.15 
6.87 
5.23 
5.03 
4.20 
