202 
D.J. Kitchener, N. Cooper, I. Maryanto 
Table 4 Canonical Variate Function coefficients for the 
three island groups in the Lesser Sundas; 
Alor; Wetar and Yamdena; The NSW 
specimen was unallocated, (see text for 
clarification). Standardised values followed by 
(in brackets) unstandardised values for (a) five 
skull and dental characters and (b) five 
external characters. For explanation of 
character codes see Materials and Methods 
section. 
Table 4a 
Character 
Function 1 
Function 2 
M 3 M 3 W 
0.7386 (5.6645) 
0.1247 (0.9561) 
M 2 B 
-0.5573 (-11.4639) 
0.6636 (13.6526) 
AOB 
-0.6868 (-5.3612) 
0.3379 (2.6373) 
MFB 
0.5213 (5.2694) 
0.0790 (0.7985) 
i,m 3 
0.3722 (2.5111) 
0.3305 (2.2301) 
Constant 
-26.4963 
-61.3732 
Variance 
Explained (%) 
65.1 
34.9 
Table 4b 
Character 
Function 1 
Function 2 
EL 
0.6592 (1.2682) 
-0.0433 (-0.0833) 
FA 
-0.1511 (-0.1473) 
1.0635 (1.0371) 
TV 
0.3056 (0.1069) 
0.3314 (0.1160) 
D4P1 
0.3745 (0.7524) 
-0.4339 (-0.8719) 
PES 
0.3451 (0.7368) 
-0.1338 (-0.2857) 
Constant 
-36.7853 
-34.2812 
Variance 
Explained (%) 
61.7 
38.3 
38.2 (35.4—41.0) 37; tibia length 19.0 (16.1-20.6) 71 v. 
16.7 (15.6-18.1) 37; and digit 3 phalanx 1 length 
17.8 (15.5-20.2) 71 v. 14.2 (12.7-15.7) 37. Forearm 
length and digit 3 phalanx 1 length both larger 
relative to digit 5 phalanx 2 length (Figures 8 a,b). 
Myotis adversus differs from Myotis macropus in 
having skull, dental and dentary measurements 
that average smaller (except for rostrum length, 
least interorbital breadth and mesopterygoid fossa 
breadth). For example, greatest skull length 16.55 
(15.72-17.20) 73 v. 16.75 (16.18-17.76) 15; brain case 
breadth 8.11 (7.70-8.40) 73 v. 8.36 (8.14-8.70) 15; 
zygomatic width 10.40 (9.60-10.75) 63 v. 10.70 
(10.20-11.42) 12; greatest skull length generally 
larger relative to both condylobasal length and 
braincase breadth (Figures 6 a,b). It averages 
smaller in all external measurements (except digit 
5 phalanx 2), and digit 3 phalanx 1 is of larger 
absolute size. For example, ear length 16.7 (14.1— 
j8.2) 71 v. 15.2 (14.3-15.8) 13; tibia length 19.0 
(16.1-20.6) 71 v. 17.5 (16.5-18.3) 13; forearm length 
42.6 (38.0-45.2) 71 v. 40.3 (38.2-41.8) 13 and digit 3 
phalanx 1 17.8 (15.5-20.2) 71 v. 14.8 (13.7-15.8) 13. 
Myotis adversus adversus (Horsfield, 1824) 
Vespertilio adversus Horsfield, 1824: 2 unnumbered 
pages 
Syntypes 
Include Natural History Museum, London, No. 
79.11.21.123; adult female; carcase in alcohol, skull 
separate. 
Type locality 
Java Island. 
Diagnosis 
Myotis adversus adversus differs from Myotis 
adversus tanimbarensis subsp. nov. in having skull, 
dental and dentary measurements averaging 
larger, except anteorbital foramen width (Table 1). 
For example, greatest skull length 16.58 (15.79- 
17.13) 46 v. 16.14 (15.72-16.66) 8; zygomatic width 
10.43 (9.81-10.75) 45 v. 10.07 (9.60-10.30) 3; I'M 3 
length 7.34 (7.01-7.61) 46 v. 7.34 (7.03-7.37) 8. M 3 M 3 
breadth generally larger relative to anteorbital 
foramen width (Figure 9). It also differs in having 
all external measurements averaging larger. For 
example, ear length 17.1 (15.5-18.2) 46 v. 15.6 (15.2- 
16.1) 7; forearm length 42.4 (40.1-44.4) 46 v. 41.2 
(39.9-43.2) 7 digit 5 metacarpal length 39.1 (36.0— 
41.8) 46 v. 36.7 (35.1-39.3) 7. Ear length larger 
relative to pes length (Figure 10). 
Myotis adversus adversus differs from Myotis 
adversus wetarensis subsp. nov. in having a 
generally longer ear 17.1 (15.5-18.2) 46 v. 16.4 
(15.8-17.4) 17; generally shorter tibia 19.0 (17.4- 
20.0) 46 v. 19.5 (18.7-20.6) 17; forearm length 42.4 
Table 5 Canonical Variate Function coefficients for the 
two locality groups: Western Australia and 
Queensland/Northern Territory (Solomon 
Island, Papua New Guinea and Seram 
specimens were unallocated in the analysis). 
Standardised values followed by (in brackets) 
unstandardised values for (a) five skull, 
dentary and dental characters and (b) five 
external characters. For explanation of 
character codes see Materials and Methods 
section. 
Table 5a 
Character 
Function 1 
AOB 
1.0628 
(12.4574) 
DL 
0.9365 
(4.3094) 
C'C'W 
-0.5684 
(-6.2501) 
M 2 B 
-0.4277 
(-5.6417) 
Constant 
-64.8959 
Table 5b 
Character 
Function 1 
EL 
1.0160 
(2.3247) 
FA 
-1.3872 
(-1.3330) 
D5M 
0.8001 
(0.8595) 
D4P2 
0.5285 
(0.7537) 
Constant 
-21.4448 
