Myotis adversus comple: 
205 
Anteorbital foramen width 
Figure 9 Plot of outside M 3 M 3 width versus anteorbital 
foramen width for male and female adult 
Myotis adversus adversus, M. a. tanimbarensis 
subsp. nov. and M. a. wetarensis subsp. nov. 
Locality codes as for Figure 2. 
Pes length 
Figure 10 Plot of ear length versus pes length for male 
and female adult Myotis a. adversus, M. a. 
tanimbarensis subsp. nov. and M. a. 
wetarensis subsp. nov. Locality codes as for 
Figure 2. 
Type 1 Type 2 
caudal cranial lateral 
Type 3 
(5 
Figure 11 Baculum types: type 1 - Myotis moluccarum 
and M. macropus; types 2 and 3 - M. 
adversus. 
Penis and baculum 
Penis ca 3.5 long; glans penis a simple pear shape 
with ventral urethral slit. Baculum with base 
bifurcated to lesser or greater extent with distal 
end narrowing (types 1 or 2, Figure 11). Maximum 
length (mean, range, N) = 0.70 (0.60-0.83) 5. 
Distribution 
Peninsular Malaysia (?); Java; Kangean Island; 
Nusa Penida Island and Nusa Tenggara (Sumbawa 
Island; Moyo Island; Flores Island; Lembata Island; 
Pantar Island; Alor Island; Timor Island and Savu 
Island). 
Myotis adversus tanimbarensis subsp. nov. 
Kitchener 
Holotype 
Museum Zoologicum Bogoriense (MZB) No. 
15906; Western Australian Museum (WAM) field 
No. M43583; adult male, skull separate, carcase 
fixed in 10% formalin, preserved in 75% ethanol; 
liver stored at Western Australian Museum in 
ultrafreeze refrigerator; baculum separate; weight 
9 gm; collected by D.J. Kitchener, R.A. How and I. 
Maryanto on 16 April 1993. 
Type locality 
7 km N. Saumlaki, Yamdena Island, Tanimbar 
Islands, Maluku Tenggara, Indonesia (7°54'00"S, 
13°20'00"E); from large cave lit by large entrance, 
with deep well to sea. 
Paratypes 
Listed in 'Specimens Examined' section. 
Diagnosis 
Myotis adversus tanimbarensis differs from Myotis 
adversus adversus as detailed in the diagnosis of the 
latter subspecies. 
