206 
D.J. Kitchener, N. Cooper, I. Maryanto 
It differs from Myotis adversus wetarensis in 
averaging smaller in all skull, dentary and dental 
measurements, except braincase breadth. For 
example, greatest skull length 16.14 (15.72-16.66) 8 
v. 16.71 (16.34-17.20) 18; zygomatic width 10.07 
(9.60-10.30) 3 v. 10.43 (10.21-10.62) 14; I'M 3 length 
7.34 (7.03-7.37) 8 v. 7.45 (7.22-7.75) 18; M 3 M 3 width 
smaller relative to anteorbital breadth (Figure 9). It 
also differs in having all external measurements 
smaller. For example, ear length 15.6 (15.2-16.1) 7 
v. 16.4 (15.8-17.4) 17; tibia length 18.5 (18.5-19.3) 
7v. 19.5 (18.7-20.6) 7; forearm length 41.2 (39.9- 
43.2) 7 v. 43.8 (42.3—45.2) 17; digit 5 metacarpal 
length 36.7 (35.1-39.3) 7 v. 39.6 (38.0-40.7) 17. Ear 
length shorter relative to pes length (Figure 10). 
Description 
The morphology of M. a. tanimbarensis is as 
described earlier for M. a. adversus except for 
differences noted in the earlier diagnosis and as 
follows: M. a. tanimbarensis has the skull at the 
junction of the parietal and frontal regions more 
inflated. The baculum of the holotype MZB 15906 
was a type 3 (Figure 11) and had a greatest length 
of 0.71. The dorsal pelage Dusky Brown; ventral 
surface basal hairs Dusky Brown tipped with Pale 
Neutral Gray. Patagia and ears Dusky Brown. 
Distribution 
Known only from Yamdena Island, Tanimbar 
Group, Maluku Tenggara, Indonesia. 
Etymology 
Named after the Tanimbar Islands. 
Myotis adversus wetarensis subsp. nov. 
Kitchener 
Holotype 
Museum Zoologicum Bogoriense (MZB) No. 
15907; Western Australian Museum (WAM) field 
No. M44690; adult female, skull separate, carcase 
fixed in 10% formalin, preserved in 75% ethanol, 
weight 13.2 gm; collected by D.J. Kitchener and 
R.A. How on 23 September. 
Type locality 
2 km E. Ipokil, Wetar Island, Maluku Tenggara, 
Indonesia, (7°45'00"S, 128°48'20"E) from a large 
sunken cave. 
Paratypes 
Listed in 'Specimens Examined' section. 
Diagnosis 
Myotis adversus wetarensis differs from Myotis 
adversus adversus and Myotis adversus tanimbarensis 
as detailed in those earlier diagnoses. 
Description 
The morphology of M. a. wetarensis is as 
described earlier for M. a. adversus except for 
differences noted in the earlier diagnosis. The 
baculum of WAM 44706 is a type 3 (Figure 11) 
with maximum length 0.71. The dorsal pelage Dark 
Grayish Brown; ventral surface with basal hairs 
Dark Grayish Brown tipped with White; patagium 
Fuscous. 
Distribution 
Known only from Wetar Island, Maluku 
Tenggara, Indonesia. 
Etymology 
Named after Wetar Island. 
Myotis adversus subsp. indet. 
Remarks 
The single adult male specimen from New South 
Wales (AM 13250) was classified by DFA to Myotis 
adversus tanimbarensis. While it is closest 
morphologically to that subspecies, it differs from 
it in having a skull with the cranium less inflated 
immediately posterior of the interorbital 
constriction; longer bulla 3.13 v. 2.96 (2.72-2.98) 
and generally smaller externally (see Table 1). For 
example, forearm length 38.0 v. 41.2 (39.9-43.2); ear 
length 14.1 v. 15.6 (15.2-16.1) 7 and tibia length 
16.1 v. 18.5 (18.0-19.3) 7. It also has a baculum that 
is more broadly spatulate (type 1, Figure 11). 
Myotis tnoluccarum (Thomas, 1915) 
Leuconoe moluccarum Thomas, 1915:170-172. 
Diagnosis 
Myotis moluccarum differs from Myotis adversus 
as detailed in the earlier diagnoses of the latter 
species. 
It differs from Myotis macropus in averaging 
smaller for all skull, dentary and dental 
measurements. For example greatest skull length 
15.64 (14.98-16.37) 37 v. 16.75 (16.18-17.76) 15; 
braincase breadth 7.81 (7.48-8.19) 37 v. 8.36 (8,14- 
8.70) 15; zygomatic width 10.06 (9.50-10.76) 34 v. 
10.70 (10.20-11.42) 12; M 3 M 3 width 6.41 (6.19-6.76) 
37 v. 6.94 (6.64-7.37) 15; I'M 3 length 6.92 (6.68-7.32) 
37 v. 7.54 (7.13-7.93) 15. M 3 M 3 width smaller 
relative to C'C 1 width (Figure 12). It also averages 
smaller in all external measurements. For example, 
tibia length 16.7 (15.6-18.1) 37 v. 17.5 (16.5-18.3) 13; 
forearm length 38.2 (35.4—41.0) 37 v. 40.3 (38.2-41.8) 
13; digit 3 metacarpal 37.2 (34.0-39.1) 37 v. 39.4 
(37.2-41.1) 13. Forearm length generally longer 
relative to digit 5 phalanx 2 length (Figure 8a). 
