Outside M 3 M 3 width 
Myotis adversus complex 
207 
7.5 
6.CH- -- •-1-- - 1--- 
4.0 4.5 5.0 5.5 
Outside C'C 1 width 
Figure 12 Plot of outer M 3 M 3 width versus outer CC 1 
width for Myotis moluccarum and M. 
macropus. Locality codes as for Figure 2. 
Myotis moluccarum moluccarum (Thomas, 1915) 
Leuconoe moluccarum Thomas, 1915: 
Holotype 
British Museum No. 10.3.1.29 (original number 
854); adult male; skin and skull separate; collected 
by W. Stalker in July 1909. 
Type locality 
Ara, Kei (= Kai) Islands, Maluku Tenggara, 
Indonesia. 
Specimens examined 
See 'Specimens Examined' section. 
Diagnosis 
[Our measurements of AM 23420 and SAM 
21781, Solomon Is are followed, where available (in 
brackets) by the mean and range of the holotype 
measurements of solormmis from Troughton (1929) 
and five specimens listed in Phillips and Bimey 
(1968)]. 
Myotis moluccarum moluccarum differs from 
Myotis moluccarum richardsi subsp. nov. in 
averaging smaller in all skull, dentary and dental 
measurements. For example, greatest skull length 
15.48 (14.98-15.82) 22 v. 15.95 (15.39-16.37) 13; 
braincase breadth 7.70 (7.48-7.86) 22 v. 8.01 (7.85- 
8.19) 13; zygomatic width 9.92 (9.50-10.28) 22 v. 
10.32 (9.74-10.76) 13; I'M 1 length 6.85 (6.68-7.06) 22 
v. 7.03 (6.74-7.32) 13; braincase breadth narrower 
than OO width (Figure 13). Ear generally longer 
relative to digit 5 metacarpal length (Figure 14). 
It differs from the Solomon Island form of Myotis 
moluccarum in having both a larger anteorbital 
foramen width 4.16 (4.00-4.32) 22 v. 3.89-3.93 and 
C‘C‘ width 4.50 (4.40-4.68) 22 v. 4.22-4.34 [4 4(4.3- 
4.4)]. CC 1 width larger relative to braincase 
breadth (Figure 13). Forearm generally shorter 37.6 
(35.4-39.7) 22 V. 38.8-40.4 [39.6 (38.5-41.3] and a 
paler ventral pelage. 
Description 
The morphology of M.m. moluccarum is as 
described earlier for M. a. adversus, except for 
differences noted in the earlier diagnosis, and as 
follows: the median postpalatal spine projects 
further posteriorly, is broader, and projects less 
ventrally; mesopterygoid fossa broad with broad 
external ventral flanges on the pterygoid processes; 
such that the sphenorbital fissure is almost 
obscured from the ventral view; and the P 3 is 
usually well intruded from tire tooth row such that 
P 1 and P 1 cingulum are often in contact. 
Baculum with base slightly broadened and distal 
shaft spatulate (type 1, Figure 11), greatest length 
0.71 (0.66-0.78) 3. 
Dorsal pelage Burnt Umber; ventral surface basal 
Outside C'C 1 width 
Figure 13 Plot of braincase breadth versus outer CC 1 
width for Myotis moluccarum moluccarum, M. 
moluccarum (Solomon Is) and M. m. richardsi 
subsp. nov. Locality codes as for Figure 2. 
