210 
D.J. Kitchener, N. Cooper, I. Maryanto 
the Kai Islands, Seram (and possibly to Peleng Is¬ 
land, Sulawesi and Irian Jaya - see Hill (1983:160)), 
New Guinea, the Solomon Islands and northern 
Australia. 
Hill (/oc. tit.) notes that M. a. carimatae from 
Borneo and Sumatra is very similar to M. 
moluccarum but notes that "only limited material is 
available for comparison". We have not examined 
that material. 
The distribution of the Myotis adversus complex 
in eastern Indonesia presents a pattern of 
morphology somewhat different again from those 
recently reported for other bat species in that 
region. For example, the form of Hipposideros 
diadema from Java ( nobilis ) reaches Lombok Island, 
where it interfaces sharply across the sea strait of 
Sumbawa with the eastern form of H. diadema 
(diadema). Rhinolophus simplex and Hipposideros 
sumbae. Lesser Sunda endemic species, are widely 
distributed throughout the Inner Banda Arc but 
differentiate into identifiable morphological forms 
on the outer Banda Islands. The pteropodid 
Pteropus lombocensis, endemic to the Lesser Sunda 
Islands, has a distinct western form (Lombok and 
Sumbawa Is) and an eastern form which occurs as 
far east as Alor Island; this eastern form shows 
some further morphological variation. The 
pteropodid Aethalops alecto has a wide distribution 
on the mountain tops of Sumatra, Java, Bali and 
Lombok Islands. However, on these latter two 
islands it has differentiated morphologically, 
sufficient to warrant subspecific status (Kitchener 
and Maryanto 1993, Kitchener el. al 1992, 1993 a-d, 
1995 a,b). Other studies, by us and our colleagues 
A. Suyanto and Maharadatunkamsi (unpublished 
data) on Scotophilus kuhlii, Hipposideros ater, 
Rhinolophus affinis, Eonycteris spelaea and 
Macroglossus minimus, also reveal a variety of 
distributional patterns of subspecies of chiroptera 
in Eastern Indonesia which no doubt reflect current 
geography, recent historic events and current 
climatic patterns in tire region. Clearly, however, 
there is a complex interaction between all these 
factors which has resulted in this mosaic of 
intraspecific morphological differentiation. 
SPECIMENS EXAMINED 
Myotis adversus adversus 
Java: Locality unknown, BMNH (401 405) (d,2). 
Kangean Island: Central region, 115°20'S, 6°55'E, 
MZB13349 (Id) 
Nusa Penida: Karangsari, 8°42'S, 115°35'E, WAM 
M (39627-839663,39674, 39680,39682) (2 d d, 4 $ 2). 
Sumbawa Island: Desa Sangeang, 8°18'S, 118°56'E, 
WAM M (31541, 31546, 31554-5, 31567) (Id, 4$ 2). 
Moyo Island: Sebotok, 8°09’30"S, 117°37T5"E, 
WAM M 31907 (Id). 
Flores Island: Ratulodong, 8°11'00"S, 122°52'00"E, 
WAM M (32569, 32574, 32586) (3d d). 
Lembata Island: Merdeka Hadakewa, 8°22'S, 
123°3LE, WAM M 32358 (12). 
Pantar Island: Batu Bakalang, 8°14'S, 124°18'E, 
WAM M 37742 (Id). 
Alor Island: Kalabahi, 8°14'S, 124°32'E, WAM M 
(37523-5, 37527, 37536, 37547, 37553, 37555, 37557, 
37581, 37583-4, 37586-7, 37589-91, 37593, 37595- 
600) (8d d, 162 2). 
Timor Island: Lifuleo Oisina, 10°18'S, 123°30'E, 
WAM M 38066 (Id). 
Savu Island: Menia, 10°29'S, 121°55'E, WAM M 
(35256-8,35265) (Id, 32 2). 
Myotis adversus wetarensis subsp. nov. (all 
paratypes) 
Wetar Island: Desa Ipokil, 7°50'S, 126°16'E, WAM 
M (44686-7,44689-94,44696,44698-706)(Id, 172 2). 
Myotis adversus tanimbarensis subsp. nov. (all 
paratypes) 
Yamdena Island: Saumlaki, 7°59'S, 131°22'E, WAM 
M (43581-5, 43589, 43618-9) (7d d, 12). 
Myotis adversus subsp. indet. 
New South Wales: Rocky Creek, 28°39'S, 153°20'E, 
AM M 13250 (Id). 
Myotis moluccarum moluccarum 
Western Australia: Drysdale R. Nat. Park, 14°40'S, 
127°00’E, WAM M (14063-5, 14067-77, 14079) 
(3dd, 122 2). Mitchell Plateau, 14°30'00"S, 
125°47'20"E, WAM M (15763, 21571, 21582) (2d d, 
12). Surveyors Pool, Mitchell Plateau, 14°40’20"S, 
125°43'40"E, WAM M (21509-11) (2d d, 12). Prince 
Regent Reserve, 15°49'25"S, 125°37'03"E, WAM M 
12255 (Id). 
Myotis moluccarum richardsi subsp. nov. (all 
paratypes) 
Northern Territory: Melville Island, 11°18’S, 
130°27 E, C 953 (sex unknown). Mungejirri Yaalput 
Waterhole, 14°32'S, 135°15'E, SAM Ml810 (Id). 
Queensland: Brisbane, 27°28'S, 153°01'E, JM 2838, 
(12). Dowah Creek, JM (5000, 5003) (2 2 2). 
Hinchinbrook Island, 18°22'S, 146°15'E, JM 5335 
(12). Jerona, Ayr, 19°34'S, 147°13'E, JM (7971-2, 
7975) (Id, 22 2). Lake Barrine, 17°16'S, 145°35'E, 
AM M 4901 (12). Macleods Creek, Cooktown, 
15°26'S, 145°08'E, AM M 13317 (12). Noosa Heads, 
