226 
I. Bartsch 
29 
Figures 27-30 Syngastes dentipes sp. nov., <5: 27, urosome with P5, lateral aspect from right hand side; 28, urosome 
posterior aspect; 29, urosome with P5, anterior aspect; 30, lateral aspect. Scale lines = 50 pm. 
truncate than in female. With three free thoracic 
somites (Figure 23), the fourth somite incorporated 
into genital complex. A1 seven-segmented (Figure 
24), segments 55, 62, 30, 45,12,15, and 16 pm long. 
Three apical segments (Figure 25) more slender 
than basal segments. Aesthetascs on Al-3 and Al- 
4 4-5 pm wide, that on Al-7 shorter and more 
slender, 2 pm wide. Number of setae: 1,11,10+Ae, 
11+Ae, 1, 2, 12+Ae. Outline and setation of A2, 
Md, Mxl, Mx2 and Mxp as in female. PI - P4 
slightly shorter but otherwise similar to those of 
females. 
P5 (Figure 26) 37 pm long, undivided, though 
suture present between wide basal and elongate 
distal portion. Basal portion with one seta, distal 
portion with one basal and three apical setae. 
Genital complex of a 434 pm long paratype 135 
pm long, 155 pm high. Anterior operculum with 
long, sharply bent spiniform projection, posterior 
valve with slender posteriorly bent spine (Figures 
27, 28). Posteroventral corner of genital complex 
extending beyond anal cone. Anterior portion of 
spermatophore reservoir with pair of lamellar 
ridges enclosing a cavity (Figure 29). 
Remarks 
Syngastes dentipes is easily distinguished from 
other species on the basis of the maxilliped: the 
wide endopodal segment has a heavily sclerotised 
cylindrical appendage (instead of a slightly 
sclerotised semi-spherical or bud-shaped tubercle) 
and a very elongate and slender tongue-like 
process; the claw has two rows of teeth. S. dentipes 
is not closely related to any of the species described 
so far, the female A1 is five-segmented, the second 
maxillae are very slender and P4-enp bears asingle seta. 
All tegastids are strongly sclerotised. The cephal¬ 
ic shield in S. dentipes, as in other Syngastes, is ven- 
trally prolonged so that the maxillipeds can be hid¬ 
den between these valves. In both females and 
males of S. dentipes, the ventral margin of the en¬ 
larged genital segment is armed with a ridge en¬ 
closing a narrow groove. In a strongly bent animal, 
this groove can enclose the posteroventral margin 
of the cephalosome (Figure 30). The groove on P4- 
endl seems to be fitted to enclose the inner edge of 
exp2. The small epimeral lamellae on the 
thoracomeres can protect the articulation between 
bases and exopods of the legs P2 - P4. 
ACKNOWLEDGEMENTS 
The sublittoral substratum was made available to 
me by P.A. Hutchings which is gratefully 
acknowledged, and thanks also go to J.W. Reid and 
V.A. Harris for helpful comments on an earlier 
draft of this manuscript. 
REFERENCES 
Bartsch, I. (1993). Syngastes craterifer, ein neuer Tegastide 
(Harpacticoida, Copepoda) von Sudwestaustralien. 
Mitteilungen aus dem Hamburgischen Zoologischen 
Museum und Institute 90:187-195. 
Bartsch, I. (1994). Three new species of Syngastes from 
south-western Australia (Tegastidae, Harpacticoida, 
Copepoda). Spixiana 17: 161-173. 
Nicholls, A.G. (1941). Littoral Copepoda from South 
Australia (1) Harpacticoida. Records of the South 
Australian Museum 6: 381-427. 
Manuscript received 15 August 1994; accepted 13 March 
1995. 
